342 Dr. A. S. Packard, Jun., on the 
while the Chilopods are a secondary less primitive group. 
Paleontology apparently supports this view. We may now 
_ turn to the structure of the head of Chilopod Myriopoda, which 
has been fully described by Newport * and also by Memert f. 
Having already briefly described the morphology of the 
epicranium or antennal segment of Chilopods, with the labrum 
and “mandibles” (protomale =“ true maxille ” of Newport), 
which are close homologues of those of diplopod Myriopods, 
we may next take up the second pair of mouth-appendages, 
which are the morphological equivalents of the so-called 
labium of Chilognaths. These, as seen in Scolopendra, are 
very different from the so-called under lip of Chilognaths ; 
they are not united, and are separate, cylindrical, fleshy, 5- 
jointed appendages, but, as Newport states, “‘ connected trans- 
versely at their base with a pair of soft appendages (e, c) that 
are situated between them, and which, as | have already stated, 
I regard as the proper Ungua, as they form the floor of the 
entrance to the pharynx.” These 5-jointed appendages are 
Mr. Newport’s “ maxillary palpi,” his true maxille being the 
homologues of the “ mandibles ”’ of Chilognaths. 
The portion of the head of Scolopendra and other Chilopods, 
thus far considered, together with the antenne and proto- and 
deutomal, we consider as homologous with the entire head 
of Chilognaths ; the basilar segment of Newport and the two 
pairs of head-appendages have no homologues in the head of 
Chilognaths. ‘They are rather analogous to the maxillipedes 
of Crustacea, and nothing like them, speaking morphologi- 
cally, exists in other Tracheata. We therefore propose the 
term malipedes (mala, jaw, pes, foot or jaw-feet) for the fourth 
and fifth pair of cephalic appendages of Chilopoda. At the 
same time it is easy to see that they are modified feet, espe- 
cially when we examine the last pair in Scolopendra, which 
are attached to a true sternite, and see that they are directly 
homologous with the feet and sternite of the same animal. 
The first pair of malipedes are the “ labium and palpi’ of 
Newport, the “ first auxiliary lip” of Savigny. They, how- 
ever, bear little resemblance to an insect’s labium and labial 
palpi. They are separate, not coalescing in the middle, as in 
the labium of Hexapods. The so-called labial palpi are 4- 
jointed, with an accessory plate; they arise directly in front 
* “ Monograph of the Class Myriapoda, Order Chilopoda, with Obser- 
vations on the General Arrangement of the Articulata,’ by George New- 
port (Trans. Linn. Soe. xix. p. 287). 
t ‘ Myriapoda Muszei Havniensis. Bidrag til Myriapodernes Morpho- 
logi og Systematik,” ved Fr. Meinert; ‘ Naturhistorisk Tidsskrift,’ 3 R. 
7B. (isi): 
