SYSTEMATIC POSITION OF TROCHOJJENBRE.E. 151 
know, when writing my paper, that such a discovery, enabling Dr, 
Seemann to advocate tlie establishment of the Natural Order Trocho- 
dendrece^ had actually come to light in the carpological structure of 
Euptdia, This genus, established by Zuccarini upon a Japanese 
species discovered by Siebold, ('Flora Japonica,' p. 133, t, 73,) was 
referred by its author to Ulmacece, in accordance with its then known 
anthological characters; but, on account of its numerous disconnected 
carpels, it occupied in that Order an isolated position. A second 
species in ripe fruit was aftei'wards discovered by Griffilh in Assam. 
At first sight this was not identified with Eapteliay and strangely 
enough, in the preliminary arrangement of the specimens, was also 
referred to Ulma€e<^, where it remained till recently more carefully 
examined by the celebrated authors of the ' Flora Indica,' Prs. 
Hooker and Thomson. It was found that Zuccarini was wrong in 
placing Euptelia, with its large quantity of albumen and minute em- 
bryo, in Uhmcece^ and that its true relationship had to be sought for 
in the neighbourliood of Wiuterace^e. Hooker and Thomson's article 
" On the Genus Euptelia " was published in No. 28 of the 'Journal of 
the Proceedings of the Linnean Society/ which, though appearin 
o* 
early in 1864, did not reach Munich till the autumn, so that when 
writing my paper I could have no knowledge of it. 
As already mentioned, Dr. Seemaun declares Eii^telia to be closely 
allied to Trochodendron, and both genera to be the first known mem- 
bers of a new Natiu'al Order. After examining the authentic speci- 
mens of both genera existing in our Academical Herbarium, I fully 
concur in his view ; and, as the subject has been mooted, I will 
briefly refer to the Natural Order constituted by the two genera, 
and the characters in which they agree and differ. In the first in- 
stance, we have in both genera the same habit, an erect shrubby or ar- 
borescent growth; alternate, simple, penninerved, exstipulate leaves, 
with a serration, the points of which are glandulose (a peculiarity which 
Zuccarini overlooked in TrocJtodtndrou') ; leaf and flower-buds are co- 
vered with protecting scales ; there is an indeterminate (racemose or cy- 
mosc) inflorescence, and scaly bracts. With regard to the structure of 
flowers and fruit, both i^gree in the total absence of a perigoninra, in 
the indefinite number of stamens and their structure, and in the inde. 
finite number of the carpels, and also in the circumstance that they are 
arranged around the very short floral axis in a single whorl, TLey 
