90 Bulletin American Museum of Natural History. [Vol. X, 



were covered with hoofs that supported the feet. This would 

 agree with the position given them in the restoration, which coin- 

 cides with the anatomical structure of the entire hind limb." 



This clear statement of Professor Marsh's as to the position of 

 these limbs is consistent with his restoration, in which not only 

 the metapodials but the phalanges are raised from the ground, 

 and the animal is represented as walking upon the tips of its toes, 

 or terminal phalanges, the latter being supported by hoofs. The 

 morphological importance of this restoration is very great. If 

 correct it places Coryphodon among the Unguligrada, widely 

 removed from the unquestionably plantigrade Pantolambda. 

 Contrary evidence that Coryphodon, so far from being unguligrade, 

 was transitional between Q.om\)\eie. plaiiti;:;radis>/i and dii:;itigrad/sm, 

 is given below. Many other important morphological characters 

 are involved in Marsh's restoration, and are now found to be 

 incorrect. The scapula is given a trihedral form, with a very 

 broad angular infraspinatus fossa ; the pelvis is extraordinarily 

 reduced ; the limbs are elongate and, together with the above- 

 mentioned unguligrade action of the digits, elevated the body 

 very much from the ground. In proportion to the scapula, the 

 humerus, the ulna and the radius are of very great length, and 

 similarly the tibia is only slightly shorter than the femur. The 

 most important character, however, is that assigned to the 

 vertebral column, there being 19 dorsals and, as far as can be 

 determined from the drawing, 6 lumbars, or D. L. = 25, a formula 

 exceeding that of the Rhinoceros. The net result of these obser- 

 vations is to give the animal the general appearance and charac- 

 ters of a modern Perissodactyl,' with the single important excep- 

 tion of the five digits preserved in the fore and hind feet. 



It appears from our more complete material that the difference 

 between the feet was exaggerated by Osborn, as already observed 

 by Marsh. There is no doubt, however, that as seen in the 

 mounted specimen, in the forward step the calcaneum rested very 

 near the ground, being separated merely by a thick plantar pad. 

 The digits of the fore and hind feet have nearly the same relations 

 to the ground. Both feet are in a somewhat similar sta}:;e of tran- 

 sition between plantigradism and digitigradism. Pantolambda has 

 a long tuber-calcis and pes like that of the Bear. Uintatherium 



