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THE GRYLLACRIDZH AND STENOPELMATIDZE OF 
AUSTRALIA AND POLYNESIA. 
By J. G. O. Tepper, F.L.S. 
[Read September 6, 1892.] 
The GRYLLACRIDZ and STENOPELMATIDA form the two last 
families in Brunner van Wattenroyls’ system of the Locustodea. 
They resemble each other considerably in general appearance, and 
approximately in habit ; but while the former possess depressed 
tarsi without pulvilli, or sole pads, and lateral expansions to all 
the joints, the latter have compressed tarsi with usually distinct 
pulvilli and the first and second joints simple. The two families 
may be thus diagnosed :— 
GRYLLACRIDZ.—Tarsi depressed, first and second joints with 
lateral lobes separated by a furrow. Fore tibie without foramina 
and above without apical spines. Hind tibize above with single 
apical spines, below with two on each margin. 
STENOPELMATID&.—Tarsi compressed, first and second joints 
without lateral appendages, mostly provided with pulvilli. Fore 
tibiz often with foramina, above with apical spines on each 
margin. Hind tibie above with single, below with two apical 
spines on both margins. 
Although both resemble the Gryllodea, or Crickets, consider- 
ably, and have been classed with them by older authors, in- 
cluding Stal, their relationship with them is quite remote, and 
the four-jointed tarsi, the structure of the wings and the ovi- 
positor relegate them to the Locustodea, notwithstanding their 
cricket-like habits and appearance. Brunner says :—“ The first 
Gryllid undoubtedly descended from some Locustid, but the 
Stenopelmatids did not descend from that Gryllid, but were—if 
one may so express oneself—a second departure of the Locustids 
in the direction of the crickets.” 
In Brunner’s monograph are 11 genera of Gryllacride and 36 
of Stenopelmatidz described as known, the former consisting of 
132 species (Gryllacris alone of 95), and the latter of 91, whose 
general distribution will be apparent at a glance at the following 
table. 
K 
