[Vor. 11 
454 ANNALS OF THE MISSOURI BOTANICAL GARDEN 
6. THE RELATIONSHIP BETWEEN THE NUMBER OF SEEDS IN THE 
LOCULES OF THE SAME FRUIT 
The correlations between the number of flowers and fruits per 
inflorescence and the actual and relative position of the fruit on 
the inflorescence, on the one hand, and the number of seeds per 
locule, on the other, have been shown to be of a very low order 
indeed. We cannot, therefore, regard any of these earlier es- 
tablished characters, or the factors which determine them, as 
having a large influence in determining the number of seeds per 
locule. 
Having failed to locate factors of material importance in the 
characters of the inflorescence, we may inquire whether there are 
unmeasurable factors which influence all of the locules of the 
same fruit in a similar manner. 
This may be done by determining the inter-locular relationship 
for number of seeds per locule in the different series. In doing 
this, symmetrical tables are formed. The number of seeds in 
each locule is considered a first and then a second member of the 
pair in combination with the other locules of the fruit. The 
values of the correlation coefficients were checked by the use of the 
intra-class correlation formulae (Harris, 713). 
The tables of data are too voluminous for publication. 
The correlation coefficients and the regression equations are 
set forth in table xxm and show a high degree of similarity between 
numbers of seeds in the locules of the same fruit. The correla- 
tion lies between 0.70 and 0.80. Since the tables are symmet- 
rical, the correlation and the regression coefficients are identical. 
The straight lines and the empirical means for the materials for 
the 2 pairs of years are shown in fig. 13. Apparently the straight 
line represents the relationship between the observed and the 
theoretical average as well as would any curve of higher order. 
The closely contiguous position of the lines for the years shows 
the generality of the laws underlying the interdependence between 
seed number i in the 3 locules of the fruit. 
C d ions for other species are few in number. 
In Sanguinaria (Harris, '10) the correlations for number of ovules 
on the 2 placentae have been shown to be of the order r — .89 
tor — .92, while the correlations for number of seeds per locule 
