44 CALIFORNIA ACADEiMY OF SOIEKCES. 



connecting the various organs with the body-wall and require here no particular men- 

 tion. 



In tlie other clitcllar somites we find bands of transverse muscles which 

 stretch diagonally (lig. 118) across the body wall from the ventral part, almost im- 

 mediately below the ventral ganglion, to the vicinity of the outer setre. The ends of 

 these muscles penetrate between the lobes of the longitudinal muscular layer (fig. 

 118, I. III.) These muscles are not continuous through the somite but are grouped in 

 bauds. 



Ventral 'papil/ft' (figs. 120, 125 to 131). Argilophilus is readily distinguished 

 from other California Eudrilidic so far known, by its ventral papilh?, occupying the 

 ventral side of the intersegmental grooves from the spermatheca to the segmeuts next 

 posterior to the clitellum. In Arfjiloi)Jnli(s marmnrn(u>i ornntiis we meet sometimes with 

 as many as 7 or 8 pairs, while in A. m. papillifer we find as high a number in a single 

 median row, under the ventral ganglion. In no instance did I find a papilla on the 

 segment itself, all invariably occurred rising from the groove between the somites, 

 being in other words intersegmental. From cross-sections it will be seen that the 

 papilla consists of two distinct ])arts, one exterior and lateral, consisting of elongated 

 hypodei-mic supporting cells, which more or less fully enclose the interior main body 

 of the papilla. This central part consists of larger or smaller gland-like bodies, vary- 

 ing in size and number according to the size and age of the papilla, etc. In some 

 papilhie these bodies fill the larger part of the pajiilla, in others they are confined to 

 the bottom or very center of the organ. As to the nature of these organs I am, as yet, 

 gomewhat in doubt, though they certainly must be considered as sensory organs 

 of .some kind. Stained with osmic acid the bodies present in sections a darker center, 

 which ap2)ears of ganglionic nature, around which are grooped larger sacs which 

 again are composed of smaller, light-refractive granules, giving the idea of a reticu- 

 lated protoplasm. Numerous nerve fibres connect with those bodies, and evidently 

 penetrate to the gaglionic center. Long tube-like cells butt directly on these gland- 

 ulous bodies, while others grooped in bundles penetrate between them connecting with 

 nerve ganglia. The transition between the supporting cells and the drainpipe cells 

 of the papilla is sudden and not gradual. In fact the line of demarcation b(;tween the 

 two is quite prominent. The supporting cells in the young papilla, entirely enclose 

 the papilla, while in the larger and full grown papilla, they are pushed towards the 

 .side leaving the whole center to the drainpipe cells and the glands. 



When stained with safranine the nuclei of the glandular cells become prominent, 

 less so when stained with ha-matoxylon. The nuclei vary greatly in size and number. 

 In some of the glandular bodies there are from one to three nuclei, in others none. In 

 sections there are always less nuclei than divisions (cells?) apparently lying on the top of 

 the reticulated mass. The nuclei may be seen directl}' above the nerve center, but 

 generally they are found outside of it. It seems as if these glandulous bodies are 

 modified agglomerations of hypodermic glands. 



The arrangement of the glandulous bodies and the tube-like cells is very 



