﻿406 
  Mr. 
  Frederick 
  Miiir 
  on 
  the 
  

  

  long 
  flagellum 
  enters 
  the 
  long, 
  spiral 
  duct 
  of 
  the 
  sperma- 
  

   theca. 
  In 
  this 
  manner 
  a 
  complete 
  tube 
  is 
  formed 
  from 
  

   the 
  testes 
  to 
  the 
  spermatheca. 
  In 
  Anomala 
  orientaUs 
  the 
  

   sac 
  is 
  large 
  and 
  the 
  functional 
  orifice 
  is 
  large 
  and 
  without 
  

   any 
  transfer 
  apparatus 
  ; 
  it 
  is 
  not 
  brought 
  against 
  the 
  

   opening 
  of 
  the 
  duct 
  of 
  the 
  spermatheca 
  but 
  to 
  the 
  opening 
  

   of 
  the 
  bursa 
  copulatrix, 
  into 
  which 
  the 
  sperm 
  is 
  discharged. 
  

   By 
  what 
  means 
  it 
  is 
  conveyed 
  to 
  the 
  spermatheca 
  it 
  is 
  

   impossible 
  to 
  say 
  at 
  present. 
  

  

  The 
  coadaptation 
  of 
  the 
  organs 
  of 
  the 
  two 
  sexes 
  in 
  a 
  

   number 
  of 
  forms 
  examined 
  by 
  Dr. 
  Sharp 
  and 
  the 
  writer 
  is 
  

   quite 
  remarkable. 
  A 
  good 
  case 
  of 
  this 
  is 
  in 
  Neolamprima 
  

   adolphmae, 
  where 
  the 
  median 
  lobe 
  is 
  small 
  and 
  produced 
  

   into 
  a 
  very 
  slender 
  and 
  long 
  flagellum 
  * 
  ; 
  in 
  the 
  female 
  

   the 
  base 
  of 
  the 
  duct 
  leading 
  to 
  the 
  spermatheca 
  is 
  widened 
  

   and 
  shaped 
  like 
  the 
  median 
  lobe 
  ; 
  the 
  more 
  distal 
  portion 
  

   of 
  the 
  duct 
  is 
  very 
  slender 
  and 
  spiral. 
  

  

  The 
  fact 
  that 
  some 
  Coleoptera 
  are 
  known 
  to 
  make 
  

   spermatophores 
  and 
  the 
  probabihty 
  that 
  others 
  do 
  not, 
  

   suggests 
  the 
  possible 
  division 
  of 
  the 
  order 
  into 
  two 
  groups. 
  

   Whether 
  this 
  classification 
  would 
  separate 
  certain 
  families, 
  

   or 
  whether 
  it 
  would 
  cut 
  across 
  several 
  families 
  separately 
  

   it 
  is 
  impossible 
  to 
  say 
  with 
  our 
  present 
  knowledge. 
  It 
  is 
  

   highly 
  probable 
  that 
  when 
  no 
  specialised 
  sac 
  is 
  present, 
  

   especially 
  if 
  the 
  median 
  orifice 
  be 
  large, 
  spermatophores 
  

   are 
  formed 
  ; 
  when 
  a 
  sac 
  is 
  present 
  and 
  no 
  specialised 
  transfer 
  

   apparatus 
  exists 
  it 
  is 
  possible, 
  or 
  even 
  probable, 
  that 
  sperma- 
  

   tophores 
  are 
  formed 
  ; 
  but 
  when 
  a 
  highly 
  specialised 
  transfer 
  

   apparatus, 
  especially 
  if 
  it 
  be 
  a 
  flagellum, 
  exists 
  it 
  is 
  highly 
  

   probable 
  that 
  no 
  spermatophores 
  are 
  formed. 
  

  

  In 
  the 
  vast 
  majority 
  of 
  Coleoptera 
  the 
  internal 
  sac 
  is 
  the 
  

   chief 
  organ 
  of 
  copulation, 
  the 
  tegmen 
  and 
  median 
  lobe 
  

   acting 
  as 
  protectors 
  and 
  guides. 
  In 
  the 
  following 
  section 
  

   the 
  muscles 
  explain 
  the 
  protrusion 
  and 
  retraction 
  of 
  the 
  

   median 
  lobe 
  and 
  tegmen 
  and 
  the 
  retraction 
  of 
  the 
  sac, 
  but 
  

   they 
  do 
  not 
  explain 
  the 
  evagination 
  of 
  the 
  sac. 
  This 
  is 
  

   always 
  brought 
  about 
  by 
  blood 
  pressure, 
  and 
  it 
  is 
  highly 
  

   probable 
  that 
  the 
  different 
  types 
  have 
  followed 
  certain 
  

   lines 
  of 
  evolution 
  to 
  accommodate 
  the 
  different 
  development 
  

   of 
  the 
  sacs, 
  and 
  allow 
  for 
  their 
  functional 
  mechanism. 
  

  

  * 
  This 
  is 
  not 
  shoAvii 
  in 
  Sharp 
  and 
  Muir, 
  Trans. 
  Ent. 
  Soc. 
  Lond., 
  

   1912, 
  PI. 
  XLIV, 
  figs. 
  10, 
  10a, 
  as 
  the 
  authors 
  failed 
  to 
  detect 
  it 
  at 
  

   that 
  time. 
  As 
  it 
  is 
  permanently 
  everted 
  it 
  is 
  easily 
  broken 
  during 
  

   dissection. 
  

  

  