20 PROF. OWEN ON CERTAIN FOSSIL BONES. 



penultimate molar ; it there defines an anterior lobe of the crown about the one-third 

 the antero-posterior extent of the crown, by a close, straight fold of enamel penetrating 

 inwards about 2i lines. 



The anterior lobe or transverse ridge of the corresponding tooth of a Lophiodon 

 might, perhaps, present a similar appearance, if worn down to the base of the crown. 



But in the present fossil, the enamel proceeds to define a middle lobe on the outer 

 side of the crown, shorter than the foregoing, beyond which the enamel and dentine 

 are worn obliquely away to the base of the posterior fang ; the indication of the 

 middle external lobe or festoon of enamel is, however, deci.sive against its generic 

 relationship with Lophiodon. 



This indication of the arrangement of the enamel, slight as it is, reminds one of 

 that in the lower molars of the Toxodon,* and another feature of resemblance is the 

 apparent interruption of the enamel at the anterior part of the molar in the fossil. If 

 the presence of two distinct fangs in the Georgian Fossil were adduced as dis- 

 tinguishing it from the Toxodon, it might be replied that perhaps the long curved 

 rootless molars in that animal at the last period of age might, as in the Horse, acquire 

 root. 



But the triloloed character is on the outside of the molar in question, and on the 

 inside of those of the Toxodon ; in which also the middle lobe only has a coat of 

 enamel, not the anterior or posterior of the inner lobes : there is no doubt, therefore, 

 that the Georgian Pachyderm is as generically distinct from Toxodon, as it is from 

 Lophiodon; and it would seem to have diminished the interval which divides the 

 strange Gliriform Pachyderm of South America, from the more normal Tapiroid 

 forms of Pachyderms which are found fossil in the old world. 



As naturalists have accepted the latinized Indian word Tajyirus as the generic 

 name of the existing American Pachyderm, which makes the nearest approach to the 

 present remarkable fossil, they will probably sanction the application to the genus 

 and species which it represents, of the name Harlanus Americanus ;t in honour of the 

 indefatigable and accomplished naturalist by whom the fossil was first made known 

 to science. 



Royal College of Surgeons, London, July St/i, 1846. 



* Fossil Jlammalia of the Beagle; pi. 5, fig. 2. 



tThe annexed Jrawing of this fossil was made in Loudon under the supervision of Prof. Owen. 



