158 LEVINE: CYTOLOGY OF HYMENOMYCETES 
long astral rays. I find that in B. versipellis also (PL. 8, FIG. 58) 
the two secondary spindles may be at an angle of about 10° to 
the longitudinal axis of the basidium. In these division figures, 
considerable variation is found in the angle formed by a transverse 
axis of the basidium and the long axis of the spindle. 
The nuclear membranes still persist at the equatorial plate 
stage (PL. 8, FIG. 59). In a polar view of the equatorial plate at 
least four distinct chromosomes can be seen. In the diaster stage 
however they are often seen as one or two masses (PL. 6, FIG. 
28, 29). I have been able, however, to demonstrate that at the 
poles also there are more than two chromosomes, as is shown in FIG. _ 
27, PL. 6. The secondary spindles may become elongated until 
the chromosome masses reach the wall of the basidium as in the 
first division, suggesting that the kinoplasmic rays are attached to 
the basidium wall and are pulling the chromosomes and centers 
to it. 
The young daughter nuclei grow rapidly in size (PL. 6, FIG.: 
30, 31) and remain for some time attached to the basidium wall. 
They then begin to move downward toward the base of the basid- 
ium and it at once becomes evident that they are connected by 
faintly stained strands with small granules which lie on the upper 
wall of the basidium at the point from which they started (PL. 5; 
FIG. 32). The origin of these granules cannot be easily determined. 
From their size, color reactions, and position in the basidium it ap- 
pears that they probably may be the centrosomes, which became 
fixed to the wall of the basidium in the process of division. The 
position of the centrosomes on the upper part of the basidium 
wall indicates the position of the future sterigmata. 
According to Petri (1902) the strands are the stretched nuclear 
membranes. I have been able to follow the development of these 
structures. In the early stages of their movement the nuclei 
resemble the beaked nuclei (PL. 6, FIG. 33) found in the Ascomycetes 
during the process of spore formation. As the main body of the 
nucleus progresses farther from the cell wall the fibrils become 
longer and thicker and resemble in all respects the strands figured 
by Maire (1902) and Fries (1911). I believe the fibrillar strand _ ; 
possibly may be analogous to astral rays; though as Petri suggests — : 
they may be due to the pulling out of the nuclear membrane. — 
