250 BRAINERD: FOUR HYBRIDS OF VIOLA PEDATIFIDA 
seeds; seeds brown, 2 mm. long; offspring notably diversiform. 
(PLATE 15, Aa.) 
Several sowings have been made of the seeds of this hybrid; 
but the ten plants of the first brood (236), whose offspring have 
been raised for two succeeding generations, are the only ones that 
will be here discussed. The forms of leaf found in the ten plants 
and in their respective broods of offspring are presented in the 
first half of TABLE I. The second half presents the corresponding 
facts regarding sixteen offspring of brood 236, plant 3, whose leaf 
had a hybrid form, quite the same as that of the mother plant from 
Illinois. 
TABLE I 
CULTURES OF VIOLA PAPILIONACEA XPEDATIFIDA FOR FOUR GENERATIONS 
V. papili h lan f ee 
ni She SERS Aart Pie As Sixteen Fs; offspring of 236-3 
Their wu. L leaf cleft: ¥ di 
pe oie Ra Sait 1a ne 0 a 
Their offspring—Fy 
Ten Fs offspring of hybrid 
Brood | Leaf Number and | 
| Their offspring—F3 468 form Brad | form | Total 
B inprrses Ra ARSE ease Pe teh fist ar eaeae ai 
tg a I Nccndbpiy and form| ! A |Aa| a 
Brood. |... Sees ps } | 
A Aa| a No. 1} A 60 12 | *22 
| : - No. 2) Aa 606 Bt Sil bees 
No. 2 A 595° 30 30 INo. 3| Aa | 607 | 4| 6| 3 13 
No. 3, Aa ee ae 3} s2 [No 4) 4 Speen um eee. 
| 59 8|20| 8 No. 5 a 609 512785 
No. 4) Aa boy gan eae ots tee 10 |No. 6] A 610 | 20 20 
No. 6| @ | 508 | 16 | 16 INo. 7} a | Grr | [xx] a 
No. 7 A 599 16 16 |No. 8] A 612 14 4 
No. 8 Aa 600 | 1|10| 3 14 |No. 9} @ | 613 16 16 
No. 9 Aa O08 fo 2g INO. 11; Aa | 6x5 | 3} 6) 3) 32 
No. 10) Aa | 602 | 6 3. | 45 |No. 12} Aa | 616 | 3 7| 3| 13 
No. 11} A 603 | 13 13 |No. 13 GETZ | 32 | 12 
No. 12) Aa 604 | 3] 5| 2 to |No. 14| A 618 |12 | 12 
0. 15 a r2|° 12 
Foo ai 199181 33 | 389 WN 6) 48 | Geo 2] 7] 4) 
| 105 _|No. 17) @ } 621 | oud Roe 
rio tA eta t | 86 | 216 
Si ee From Aa’s.../ 15 |31|18| 64 
The cultures show that in this hybrid the Mendelian law 
controls in a general way the inheritance of leaf form, though there 
is no dominance, the hybrid leaf being intermediate between the 
leaves of the two parents. The plants that resemble V. pedatifida 
in having parted leaves always produce offspring with parted 
leaves; those that resemble V. papilionacea in having uncut 
leaves always produce offspring with uncut leaves; while those 
