328 GLEASON: STUDIES ON WEsT INDIAN VERNONIEAE 
brown and finely tomentulose; leaves numerous, crowded, firm, 
dark green, oblong to elliptic-obovate, broadest at or above the 
middle, the principal ones 2.5-4 cm. long and I-1.5 cm. wide, 
obtuse or subacute, entire, obtuse or rounded at the base, scabrel- 
late and resinous-punctate above, glabrous beneath and densely 
punctate with resinous globules and impressed black glands; 
midvein puberulent, the lateral veins inconspicuous; petioles I-2 
mm. long; inflorescence terminal, irregular in shape, consisting 
of several short, simple or sparingly branched cymes 2-4 cm. long, 
naked below, and bearing 2-8 crowded heads in a terminal sub- 
capitate cluster, or at the base of the branches; bracteal leaves 
I—3 subtending each cluster of heads, resembling the cauline in 
shape, 5-15 mm. long; heads about 8-flowered; corollas white; 
involucre campanulate, 3-4 mm. high, its scales loosely and ir- 
regularly imbricated, appressed at the base, but spreading at the 
tip, stiff and firm in texture, the outer narrowly triangular-lanceo- 
late, long-acuminate, the inner narrowly oblong-linear, tapering 
gradually to the acuminate puberulent apex; achenes thinly 
pubescent, 2 mm. long; pappus nearly white, the outer series I 
mm., the inner 4 mm. long. 
Type, Shafer 4050, from rocky river banks in the vicinity of 
Camp San Benito, Oriente, Cuba, altitude 900 m., February 24, 
1910, deposited in the Herbarium of the New York Botanical 
Garden. Other sheets in the same herbarium, all collected in the 
mountains of Oriente, are Shafer 8051, stated to be vinelike and 8 
feet high; Shafer 8216, 1.5-2 feet high; and Shafer 4446, described 
by the collector as an herb three feet high with purple flowers. 
Notwithstanding field differences in the color of flowers or texture 
of stem, all four numbers clearly belong to the same species. 
The relationship of V. segregata is puzzling. The subcapitate 
clusters clearly represent a modification of a scorpioid type, and 
most closely resemble the inflorescence of the Scorpioideae aggre- 
gatae. The involucre is quite different, however, from that of 
typical members of the group. For the present, it has been con- 
sidered advisable not to assign the species to any group. 
SPECIES-GROUP HAVANENSES 
In recent work on Vernonia, V. havanensis and V. Ottonis have 
been considered identical (Revision, 192). The large series of 
specimens now available for study permits the ready separation — 
of two species, with characters so typical that to each can be 
