377] ANOPLOCEPHALIDAZ—DOUTHITT 27 



whatever except to external form; and Galli-Valerio (1905) burdens 

 science with A. dentata, with but two lines of information as to internal 

 anatomy and these lines devoted to non-essential details. There is no 

 proof that either of these two species belongs even to the subfamily; 

 and indeed, the only important fact made known concerning A. minima, 

 that the genital pores are regularly alternate, indicates that it does not 

 belong to the genus Anoplocephala. 



If consideration be given to these half-starved species, no anatomi- 

 cal generalization concerning the genus Anoplocephala is possible. Thus, 

 "A." spatula has the testes extending across the median field; "A." 

 omphalodes has the genital pores irregularly alternate, the vaginal pore 

 posterior, and the testes sometimes extending across the median field; 

 and "A." minima has the pores regularly alternate. For about half 

 the species assigned to the genus, no statement whatever is made con- 

 cerning the uterus. There seems no alternative, then, if one would 

 attempt any generalizations, but to base them only on those species 

 whose position in this genus is assured, and to ignore seeming contra- 

 dictions coming from the inadequately described species which are 

 placed provisionally in the genus. 



The diagnosis of the genus should be changed so as to admit species 

 in which the testes extend beyond the excretory ducts on the side away 

 from the genital pore. Also, it should state that the vagina and vaginal 

 pore are ventral to the cirrus pouch. It seems probable that in all the 

 proximal end of the oviduct lies beneath the median lobe of the vitelline 

 gland ; but information is needed concerning more species on this point. 



The diagnosis of the genus Anoplocephala thus becomes as follows. 



Anoplocephalinae, with segments generally much broader than 

 long, occasionally longer than broad. A single set of reproductive or- 

 gans in each segment. Genital pores unilateral. Genital canals pass 

 on the dorsal side of the longitudinal excretory vessels and nerve. 

 External vesicula seminalis present. Testes on the side away from the 

 genital pore, sometimes extending laterad of the nerve trunk. Vagina 

 and vaginal pore ventral to the cirrus pouch. Female glands to the 

 pore side of the median field. Uterus a transversely elongated sac, with 

 pocket-like appendages, anteriorly and posteriorly. Eggs with well 

 developed pyriform apparatus. Adults in mammals. 



Type-species : Anoplocephala perfoliata Goeze 1782. 



