18G ALTERATION OF POSITION. 



less split open at the' distal end, indicating a tendency 

 towards dialysis. The pollen-bearing ovules were 

 borne on the edges of these ovaries, and presented 

 various intermediate conditions between anthers and 

 ovules, commencing at the distal extremity of the 

 carpel with a bi-lobed anther, and passing in series to 

 the base of the ovary, an antheroid body of ovule-Uke 

 form, a modified ovule containing pollen, an ovule 

 departing from a perfectly natural condition only in 

 the development of a few grains of pollen in its nucleus, 

 and, finally, a perfect, normal ovule. 



In the flowers of the Rose the stamens exhibited 

 almost every conceivable gradation between their ordi- 

 nary form and that of the carpels, while some of the 

 ovules contained pollen in greater or less abundance. 

 Speaking generally, the most common state of things 

 in these flowers was the occurrence on the throat of 

 the calyx, in the position ordinarily occupied by the 

 stamens, and sometimes mingled with those organs, of 

 twisted, ribbon-like filaments, which bore about the 

 centre one or more pendulous, anatropous ovules on 

 their margins. Immediately above the latter organs 

 were the anther-lobes, more or less perfectly developed, 

 and surmounting these a long style, terminating in a 

 fringed, funnel-shaped stigma. Sometimes the ovules 

 were perfect, at other times the nucleus protruded 

 through the foramen, while in a third set the nucleus 

 was included within the tegument, the ovules having 

 in all respects their natural external conformation, con- 

 taining, however, not only pollen-grains, but also a layer 

 of those pecuhar spheroidal cells, including a fibrous 

 deposit, which are among the normal constituents of 

 the anther. In one case, where the coat of the ovule 

 was imperfect, and allowed the nucleus to protrude, 

 the pollen was evidently contained within the central 

 mass of the structure. In this instance the fibrous cells 

 were not detected, these being only found in cases where 

 the investment of the ovule was perfect ; and hence it 

 seems likely that the fibrous cells were part of the coat 



