Oenothera Scintillans. 387 



In the summer of 1897 I selfed five of the La- 

 marckianas in this culture with their own pollen. Each 

 bore from 12-13 cm. of seed, of which some was sown 

 next year in the garden and in pans. 117 seeds germinated 

 in the garden and 1079 in the pans. There was not a 

 single example of scintillans amongst these. The major- 

 ity of them were Lamarckianas, with a considerable ad- 

 mixture of mutants. In the garden these were, 4 O. 

 ruhrinervis, 3 O. lata, 1 0. nanella, 1 O. alhida and 2 

 O. oblonga ; in the pans the only mutants were 7 ex- 

 amples of O. nanella. 



The Laniarckianas, therefore which are produced by 

 0. scintillans, have the same constancy as the original 

 Lamarckiana, that is to say their grand-parents, but also 

 exhibit the same degree of mutability. 



The point is that a continued segregation into La- 

 marckiana, scintillans and oblonga is not witnessed in 

 the seedlings from the Laniarckianas extracted from scin- 

 tillans as it was in the original scintillans. 



Of the mutants mentioned O. rubrinervis, 0. lata and 

 O. nanella flowered the same summer. 



The very important question now presented itself, 

 how the scintillans plants in this generation behaved on 

 self-fertilization. To answer this question I enclosed 

 over 50 plants of the 1898 culture in bags, harvested 

 their seed separately and sowed it. All the seedlings 



