Tricotylous Races Do Not Arise by Selection. 409 



in the best cases 2.4 to 4.1 and 5.4%. The counts were 

 made on 700, 800 and 2000 seedHngs. These three plants 

 had, however, decussate leaves on their main stems, and 

 this shows that the disposition of leaves on the stem is 

 of subordinate value in selection. In the later genera- 

 tions I have always selected the ternary individuals where 

 possible, but have not found that they are any better than 

 the others. 



In the summer of 1894 tricotylous seedlings of the 

 seed-parent w^ith 5.4% were planted out. I saved the 

 seeds of 25 plants separately, and in the following spring 

 examined from 300 to 500 seedlings in each crop. The 

 values were essentially the same as in the previous year, 

 and varied between 0.5% and 5.5% with a mean of 2% : 

 and the five best gave 3.6 — 3.8 — 4.0 — 4.2 and 5.5%; 

 the three last values are based on counts of 1000, 1500 and 

 2000 seedlings. It was only the offspring of these parents 

 that were planted out in 1895, and only those with three 

 cotyledons and with a first whorl of three leaves (Fig. 

 73, p. 366). 



By planting them out in three groups in the following 

 spring, a selection was made possible which related to the 

 grandparents. Those with 4.0 to 4.2% gave offspring 

 with 1.3 to 5% with a mean of 3% ; (from 12 parents 

 and from counts of groups of 300 seedlings). The 

 grandparent with 5.5% proved to be considerably better. 

 Its twelve offspring had 2 to 8%, with a mean of 4.5%. 

 Seedlings of this group only were planted out in 1896 

 and only tricotylous specimens with ternary lower whorls, 

 from parents with 6 — 6 and 8%. They were in all 72 

 strong plants each of which gave a sufficient harvest for 

 the calculation of their hereditary values in groups of 

 300 seedlings. The parent with 8% gave 2 to 15%, with 



