J. M. Macfarlane. — Sarraceniaccae. 



more striking were the leaves on a semi-aquatic plant of the latter, gathered by the 

 writer during winter near Ochesee, Fla. One leaf was %\ cm long, the pitcher was 

 a long narrow tube 15 cm X 0,5 cm, but the wing was M cm long, and at its widest 

 part 6 cm deep. A double set of vascular bundles runs into the wing from varying 

 heights of the midrib, and distribute themselves over either face of it as reticulate veins. 



The lid varies in size and morphological relation in the different species. In 

 S. rubra and to less degree in S. Sledgei and S. Drummondii, it is slightly but 

 more or less conoave , and bf ovate or nearly circular outline. Here it is separated 

 from the pitcher mouth by a constricted neck, into which pass the longitudinally dis- 

 posed pitcher veins , that have ascended dorsally, or that have curved round the mouth 

 and reformed by fusing, hefore passing up into the lid. Traversing the median dorsal 

 part of the tube , and prolonged into the lid is a 

 vein that ends in the organic apex of the lid as 

 an acuminate process [S. rubra, S. Sledgei), or that 

 may become excurrent as a free process that is 

 abundantly loaded with honey glands, as in some 

 leaves of S. flava. When such is formed, the 

 laminar substance of the lid may grow forward 

 as a slight peltation beyond the excurrent point. 

 Similar growth, to a more pronounced degree, causes 

 a peltate fusion in S, psittacina (Fig. 3) beneath 

 and in front of the beak-like apex of the vein, 

 which can scarcely be said to be excurrent, though 

 strongly protruding. The reunited veins from the 

 pitcher wall also run up into 'the lid, parallel to the 

 median one, with which they form anastomoses, 

 and then distribute radiating lateral veins into the 

 laminar halves of the lid. Morphologically therefore 

 these should be regarded as laminar lobes, developed 

 on either side of the prolonged and flattened neck 

 of the midrib (Fig. \ Cf). In S. minor the lid 

 scarcely differs from that of the seedling, except 

 that it becomes more arched. 



Embryologically each pitchered leaf arises near 

 the growing apex of the rhizome as a conical papilla 

 with expanded base. Along its ventral aspect a median 

 ridge arises, and above this, in line with it, an oval 

 depression appears, the rudiment of the future cavity. 

 The writer regards the ridge as the fundament of the 

 lamina. The future pitcher thus arises as a ventral 



Fig. 4. Sarracenia purpurea L. A Ha- 

 bit. ; several leaves and a part [of 

 the inflorescence removed. B Flower- 

 bud, showing the calyculate bractlets. 

 (Cop. from A. Gray, Gen. fl. amer. 

 ill. t. 45 and Engler-Prantl , Pflzfam. 

 III. 2, p. 24 6 fig. U7.) 



median depression of the midrib at the apex of the 

 fused laminar halves, and slightly beneath the organic apex of the leaf. As growth 

 proceeds the steadily deepening depression occupies an increasingly large area of the medul- 

 lary tissue of the rudiment. The median ridge or crest continues to grow out as the future 

 laminar wing, while between this and the sheathing base a solid cylindrical part lengthens to 

 form the upper area of the petiole. The apex of the rudiment, at a later period, lengthens 

 into a terminal process, on whose edges arise two closely involute outgrowths, the 

 laminar halves of the lid. When mature the entire leaf thus consists of (a) a petiolar 

 portion, and (b) a pitchered laminar portion. The former is made up of a broadly 

 sheathing base, whose wings higher up converge into a faint median line on the ventral 

 face, and of an upper solid cylindrical portion. The latter (b) consists of the greatly 

 enlarged and pitchered midrib, that bears on its ventral face two fused inferior lobes 

 of the lamina as a median wing. Goebel is evidently in error when he considers that 

 the occasional bilobed or two-ridged condition, sometimes observed in the wing, is to 



