XII AFFINITIES OF ZEUGLODON 385 
instead of rudimentary like those of other Whales, the blow- 
hole lies more in the middle of the face. The skull, too, is 
not Whale-like in a number of other points. Thus the pre- 
maxillaries take their fair share in the outline of the upper jaw ; 
and, furthermore, bear the incisor teeth. The parietals meet 
above in a crest and are not excluded from the roof of the 
skull. The vertebrae of the neck are in no way shortened ; 
neither are they fused together. The ribs aré double-headed, and 
the sternum is made up of several pieces. Some naturalists, 
particularly Professor D’Arcy Thompson," have assigned a relation- 
ship to the Seals to these ancient Cetacea; but others” have 
disputed this view chiefly on the grounds that the characters 
which appear to be Seal-like are simply characters which are 
generalised and so far at most not Whale-like. Thus the long 
neck and the serrated character of the teeth may be accepted as 
Seal-like on the one hand; but on the other, a simple serrated 
tooth and a long neck are not by any means features of organisa- 
tion which we should consider out of the way in an ancient form 
of Cetacean which probably preyed upon fish. The humerus of 
Zeuglodon, according to Mr. Lydekker, puts out of court any 
possible near relationship to the Seals. But the matter under 
dispute can be further studied by reference to the three memoirs 
quoted below. 
1 Thompson, Studies Mus. Dundee, i. 1890 ; and C. R. Congres de Zoologie, 1889, 
p. 225. 
2 Lydekker, Proc. Zool. Soc. 1892, p. 560. 
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