506 ESCALLONIACEAE 



1. HYDRANGEA L. 



Shrubs or trees. Leaves opposite, without stipules : blades entire, toothed, or lobed. 

 Flowers perfect, except a few conspicuous ones around the edge of the corymb or thyrse. 

 Sepals united into a truncate or 4-5-toothed border, or in sterile flowers of 4-5 spreading 

 petal-like lobes. Petals 4-5, valvate. Stamens 8-10, inserted at the base of an epigynous 

 disk : filaments filiform. Ovary inferior, completely or partially 2-4-celled : styles 2-4, 

 distinct or slightly united at the base. Ovules numerous, borne on axile placentae. 

 Capsule 2-4-celled, membranous, opening at the top between the styles. Seeds numerous, 

 minute. Testa membranous, reticulated. The plants flower during the spring and summer. 



Leaf-blades merely toothed : inflorescence corymbose. 



Leaf-blades glabrous or inconspicuously pubescent beneath. 1. H. arborescent. 

 Leaf-blades conspicuously and copiously pubescent beneath. 



Leaf-blades membranous, light gray beneath. 2. H. cinerea. 



Leaf-blades leathery, bright white beneath. 3. H. radiata. 



Leaf-blades deeply lobed : inflorescence thyreoid. 4. H. quercifolia. 



1. Hydrangea arborescens L. A branching, sometimes straggling shrub 1-3 in. tall. 

 Leaf -blades ovate, oval or elliptic, 0.5-2 dm. long, short-acuminate, serrate, acute, rounded 

 or cordate at the base, glabrous or sometimes minutely pubescent beneath ; petioles shorter 

 than the blades : corymbs 5-20 cm. broad, usually with few sterile ray -flowers or these 

 wanting, commonly pubescent: hypanthium campanulate, 1-1. 5 mm. broa'd : sepals minute, 

 triangular : petals deciduous, concave, ovate, about 1 mm. long : stamens conspicuously 

 exserted : stigmas reddish : capsules 2 mm. broad, 8-ribbed, broader than high, each tipped 

 with the 2 spreading styles. 



In woods or along streams, New York to Iowa, Florida and Louisiana. 



2. Hydrangea cinerea Small. A spreading shrub, 1-2 m. tall, with deep green 

 foliage. Leaf-blades thinnish, oval, elliptic, narrowly ovate or orbicular-ovate, 6-15 cm. 

 long, acuminate, serrate, obtuse, rounded or cordate at the base, bright green above, gray- 

 tomentose and not reticulated beneath ; petioles somewhat more than one-half as long as 

 the blades or shorter : corymbs 5-15 cm. broad, rather round-topped : sterile ray-flowers 

 commonly present : hypanthium campanulate, ribbed : sepals triangular, acute : petals 5, 

 ovate, boat-shaped, 1.5 mm. long, hooded at the apex : stamens conspicuously exserted: 

 capsules urn-shaped, about 2 mm. in diameter, usually higher than broad, strongly ribbed, 

 each tipped with the 2-3 spreading styles. 



On the slopes of the Alleghenies, North Carolina, Tennessee, Georgia and Alabama. 



3. Hydrangea radiata Walt. A branching shrub, 1-2.5 m. tall. Leaf -blades leathery, 

 ovate, varying to oval or ovate-lanceolate, 6-12 cm. long, acuminate, serrate or c remit c- 

 serrate, acute, rounded or subcordate at the base, bright green and glabrous above, densely 

 silvery white-tomentose and reticulated beneath ; petioles about \ as long as the blades or 

 shorter : corymb round-topped, 4-12 cm. broad : sterile flowers always present : hypan- 

 thium urn-shaped, ribbed : sepals triangular or usually somewhat acuminate : petals 5, 

 ovate, concave, nearly 1 mm. long, apiculate : stamens much exserted : capsules urn- 

 shaped, 2 mm. broad, strongly ribbed, shorter than their pedicels, each tipped with the 2 

 spreading swollen styles. 



On rocky slopes, North and South Carolina. 



4. Hydrangea quercifolia Bartr. A stout shrub, 1-2 m. tall, with densely tomentose 

 twigs and branches. Leaf-blades oval or suborbicular in outline, usually o-lobed, 6-20 

 cm. long, tomentose beneath, the lobes serrate and often slightly lobed ; petioles %-?, as 

 long as the blades : panicles 1-2 dm. long : sterile flowers numerous, white, turning purple : 

 hypanthium urn-shaped, longer than the pedicel : sepals ovate, about ^ as long as the 

 hypanthium: petals 5, oblong, 1.5 mm. long, slightly concave: stamens conspicuously 

 exserted : capsules urn-shaped, 2.5 mm. in diameter, ribbed, each tipped with the spreading 

 much enlarged styles. 



On river banks, middle Georgia to Florida and Mississippi. GRAY-BEARD. OLD MAN'S-BEARD. 



2. PHILADELPHIA L. 1 



Shrubs, with branching stems. Leaves opposite, deciduous : blades simple, petioled : 

 stipules wanting. Flowers solitary, corymbose or racemose, either terminal or axillary. 

 Hypanthium turbinate, adnate with the ovary. Sepals 4 or 5, persistent. Corolla white 

 or cream -colored. Petals 4 or 5, obovate or rounded, convolute in the bud. Stamens 

 numerous, inserted on the disk : filaments slender. Styles 3-5, united, at least at the base : 



1 Contributed by Mr. C. D. Beadle. 



