THE CAT TRIBE 353 



semblage of Mammals which is so generally carnivorous as the present one, and 

 accordingly the name by which it is designated is the most appropriate that could 

 have been selected. 



There has been a considerable amount of more or less unprofitable discus- 

 sion as to whether the Carnivores, or Apes and Monkeys, are entitled to occupy the 

 highest place among Mammals. Putting man on one side, there can, however, be 

 but little doubt that, for their particular mode of life, the higher Carnivores, both 

 as regards their bodily structure and their brain power are fully as highly organized 

 as the Apes ; and to say that the one group is higher or lower than the other is thus 

 practically an impossibility. A more just view is to compare the Carnivores and 

 the Primates with two trees of different kinds, each of which has attained practi- 

 cally the same height, and bears fruit and flowers of an equally perfect development. 



Had we to deal only with the existing forms of the animal kingdom, and if the 

 seals and walruses were excluded (as is done by some zoologists) from the Car- 

 nivores, there would be no great difficulty in giving a short and concise definition 

 which would at once distinguish the order from all the others. The seals and wal- 

 ruses differ, however, so markedly in the characteristics of their teeth, as well as in 

 many other structural points, from the more typical Carnivores, while a number of 

 extinct forms appear to connect the latter on the one hand with the Insectivores, 

 and on the other with the Marsupials, that any such concise definition is impossible. 



Among the characteristics common to all Carnivores, whether terrestrial or 

 aquatic, the following are some of the most important. In all cases the toes are 

 provided with claws, which are very generally sharp and curved, with no resem- 

 blance to nails. Then, again, the number of complete toes is never less than four 

 to each foot, and is frequently five. And in no case is the first toe capable of being 

 opposed to the other digits ; so that a Carnivore can in no sense be said to have a 

 hand in the popular acceptation of that term. 



The teeth, in conformity with the flesh-eating habits of the great majority of 

 the members of the order, are generally large and well developed ; and are always 

 divisible into incisors, tusks or canines, and cheek-teeth. As a general rule, the 

 incisors are three in number on each side of both the upper and lower jaws, 

 and in no case do they exceed this number ;* while the third or outermost of these 

 three incisors is always larger than either of the others, more especially in the upper 

 jaw. The tusks are large, and adapted for seizing and retaining the prey of these 

 animals. The different families of the order show 7 a considerable diversity in the 

 form and structure of the cheek-teeth ; but, as a general rule, the more anterior 

 of these teeth have sharp and more or less compressed crowns, while very fre- 

 quently, as will be explained later on, one pair of teeth in each jaw is specially 

 modified to bite with a scissor-like action against an opposing pair in the opposite 

 jaw. Moreover, in such Carnivores as have the crowns of the molar teeth flattened 

 and expanded, these crowns are not divided into distinct portions by infoldings 

 of the enamel, as we shall find to be so frequently the case with those of the 

 Rodents. 



*The Marsupial Carnivores never have less than four pairs of incisor teeth in the upper jaw. 

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