510 



CARYOPHYLLACEAE 



Sea bluffs of Marin and Sonoma cos. 



Locs. Pt. Reyes, Davy 6876; Bodega Head, K. Brandegee. 



Var. pacifica Jepson n. comb. Much more slender and less densely leafy; 

 leaves narrower, the basal 2 to 3 inches long on petioles nearly twice as long ; 

 claws glabrous. Sea coast, central and northern California. A transition to 

 S. verecunda. 



Locs. San Francisco, K. Brandegee; s. Marin Co.; Bucksport near Eureka, Tracy 2141. 



Befs. -SILENE GRANDIS Eastw. Bull. Torr. Club, 30: 487 (1903), type loc. Bodega Pt., 

 Eastwood. Var. PACIFICA Jepson. 5. pacifica Eastw. Bot. Gaz. 41: 285 (1906), type loc. 

 Eodeo Lagoon, near Pt. Bonita, Marin Co., Eastwood. 



19. S. douglasii Hook. var. monantha Rob. Stems erect, several from the 

 loosely branching crown of a taproot, 10 to 20 inches high, the leaves chiefly 

 basal, the stem with mostly 1 or 2 remote pairs; herbage very minutely 

 pubescent or nearly glabrous, especially below ; leaves linear-lanceolate to 

 oblanceolate, tapering to both ends, 1 to 2y 2 inches long; stems (or the main 

 branches) 1 or more commonly 3 to 5- flowered; calyx oblong-cylindric, soon 

 inflated and oblong-campanulate, 6 to 7 lines long, its teeth roundish, often 

 a little constricted at base, obtuse at apex or at length with the membranous 

 margins inflexed and thus acute; corolla dull white, 5 to 8 lines broad; petal 

 blades 2-cleft, its lobes entire ; claws somewhat exserted ; scales oblong, entire ; 

 auricles obtuse or acute ; capsule elliptic or oblong, 4 to 5 lines long, included, 

 on a stipe 1% lines long. 



Sierra Nevada, 6000 to 8000 feet, from Placer Co. north to Modoc Co., 

 thence to western Siskiyou. North to Washington. 



Locs. Fallen Leaf Lake, Hall 8773; Cisco, Placer Co., Hall 8728, 8742; between Donner 

 Lake and Coldstream, Heller 6957; Tallac, El Dorado Co., C. J. Fox, Jr.; Nevada Co., Carpen- 

 ter; Lower Sardine Lake, Sierra Co., Hall $ Bdbcoclc 4491 ; Dixie Valley, Lassen Co., Baiter 

 <$ Nutting ; Mt. Bidwell, Mary H. Manning ; Highland Mine, Siskiyou Co., Butler 963. 



Specimens from Sierra, Placer and El Dorado counties in the northern Sierra Nevada are 

 very uniform in habit save in number of flowers on the flowering stems. One finds specimens 

 with the stems 3 or 5-flowered, rarely 7-flowered. Less commonly plants are found with all 

 the stems 1-flowered, but frequently plants show all these variations on a single individual. 

 In his revision of Silene, Williams, whose material was evidently scanty, disposes of S. monantha 

 Wats., the type of which is simply the 1-flowered state, in the subgenus Gastrosilene and places 

 S. douglasii Hook, (the many-flowered original form) in the subgenus Eusilene. In the absence 

 of ample material a specific unit may thus be broken and separated, by applying too rigidly 

 the characters of defined subgenera, which nevertheless may be sound in principle. 



Eefs. SILENE DOUGLASII Hook. Fl. Bor. Am. 1: 88 (1829), type spms. collected above 

 the Grand Bapids of the Columbia and on the western slope of the Eocky Mts. by Douglas. 

 Williams, Jour. Linn. Soc. Bot. 32: 143 (1896). Var. MONANTHA Bob. Proc. Am. Acad. 28: 

 145 (1893). S. monantha Wats. Proc. Am. Acad. 10: 340 (1875), type loc. Castle Eock, Cas- 

 cade Mts., Wash., Kellogg 4" Harford. S. lyallii Wats, at least as to Californian distributions. 



20. S. grayi Wats. Stems erect, caespitose, 4 to 7 inches high, arising 

 from the branching crown of a taproot; herbage finely puberulent, glandular 

 above; leaves linear to oblanceolate, 5 to 8 lines long, mostly 2 to 3 lines 

 broad, somewhat fleshy, densely crowded at base, the cauline pairs few and 

 reduced ; flowers 1 to 4 or 5, in a loose terminal cluster ; calyx purplish, broadly 

 cylindrical, soon ovoid-distended, 5 lines long, the teeth rounded ; corolla pink, 

 3 to 4 lines broad ; petal blades bifid, the segments each bearing a lateral 

 tooth ; scales lanceolate ; auricles narrow, truncate ; capsule obovoid, 4 to 5 

 lines long, the stipe almost none. 



High montane, above timber line, 7000 to 8000 feet: Mt. Shasta to ne. 

 Siskiyou, west to the Klamath Range. 



Locs. Medicine Lake Mts., M. S. Baker; Horse Camp, Mt. Shasta, Jepson; Mt. Eddy, 

 E. B. Copeland 3853; near Preston Peak, Jepson 2882. 



Eefs. SILENE GRAYI Wats. Proc. Am. Acad. 14: 291 (1879), type loc. Mt. Shasta, 

 Brewer, HooTcer $ Gray, A. S. Packard, Jr.; Bob. Bot. Gas. 16: 44, pi. 6, figs. 7, 8 (1891). 



