W. F. A. Weldon, F. Galton, E. R. Lankester 287 



mind that allowed him to co-operate effectively with other 

 men, he rendered many useful services to science. He was 

 knighted in 1909, two years before his death. By his will 

 he left a sum amounting to about 45,000 for the foundation 

 of a chair of Eugenics in the University of London, expressing 

 the wish that Karl Pearson should be the first occupant of 

 tKe chair. 



One of the rules laid down for the writers of this book 

 is that living authors should only be mentioned when their 

 work is so much interwoven with that of others whose 

 activities have been noticed that a wrong impression would 

 be created by omitting all reference to them. Professor 

 Sir E. Ray Lankester has added so much to our conceptions 

 of the morphology of the animal kingdom, so much more 

 than any other living man, that a short account of his re- 

 searches must be given. Mention must be made of his 

 investigation into the embryonic cell gland of the Mollusca, 

 his researches in the distribution of haemoglobin in the 

 Invertebrata, the wonderful way in which he, in collabora- 

 tion with one of his pupils, cleared up the structure of the 

 Lamellibranch gill, his work on the anatomy of the Limpet, 

 and the even more important series of investigations which 

 led to the assignment of Limulus to its proper position 

 amongst the Arachnids. He was the first to observe an 

 intracellular parasite (in the red corpuscle of the frog), but 

 from the scales of fossil fishes to the details of the Okapi, 

 there are few subjects in Zoology that do not owe something 

 to the investigations carried on by Lankester from 1862 

 to 1905. His name will ever be associated with the very 

 important and fundamental conception of the coelom, and 

 his views on this subject are set forth at length in Part II 

 of his Treatise on Zoology. With this theory must be 

 associated his views on Phleboedesis, a name given to the 

 theory that the lacunar blood-holding spaces forming the 

 haemocoel of the Mollusca and the Crustacea have no 

 connexion with the coelom, although they encroach in 

 certain animals on the space occupied by the coelomic 

 cavity. The discussion of how his theory differs from that 

 given in " Die Coelom Theorie " of the Hertwigs is set out 

 in the above-mentioned treatise. 



