AUTOGAMY BY THE BENDING OF THE PISTIL. 



351 



salad (Valerianella Auricula, V. carinata, &c.), and of the non-twining species of 

 Honeysuckle (Lonicera alpigena, L. nigra, L. Xylosteum) are protogynous, and the 

 style is so placed when the corolla opens that its stigma must of necessity be brush ><1 

 by insects as they make their way to the receptacle. There is evidently at this st 

 no possibility of any but cross-fertilization. Later on, when the anthers op<-n and 

 expose their pollen, the stigma is taken quite out of the way, the style becoming 

 bent downwards or to one side, so that the stigma cannot come into contact with 

 anthers of the same flowers either spontaneously or through the agency of insects. 

 Not till the flowering period is nearly over does the style return to its original 

 position; when this happens, the stigma is raised by the straightening up of the 

 style, and is pressed against the anthers, which are still coated with pollen. The 

 flowers of the Martagon Lily (Lilium Martagon) are nodding, and have their 



r 



Fig. 299. Autogamy effected by means of an inflection of the style. 



Tricyrtes pilosa in the first stage of its development. The same in the last stage of development. 



Persica in the first stage of development * The same in the last stage of development 



n f onthora hplnnirinff to the same flower Flower of Euphrasia minima in the first stage of 



witn ponen irom anuieis ueiuugiug rnat m\KA In flirm. and : the 



development, r The same in the last stage of development. All the figures slight! 

 front part of the flower is cut away 



perianth-lobes strongly reflexed. In each petal there is a groove which is roofed 

 over and closed in the middle by two coalescing rims, so that the honey 8 

 the groove can only be sucked out by insects at the two extremit.es wh. 

 groove is open. The flowers are protogynous, and, when they are newly open 

 style is straight and holds its stigma in such a position that it is sure to b 

 by insects sucking honey from the inner orifice of the groove 

 still closed at this stage, only foreign pollen can be affixed to the st.gma 

 quently dehiscence ensues, and the position of the anthers m fron of the exten 

 orifice of the groove ensures their pollen being brushed off by .nsects faying to . 

 the honey at that end, whilst the stigma is left untouched by them. In ,, ~l 



