112 



CORDAITEAE. 



a female specimen, with which all the rest agree perfectly in the structure 

 of the ovule ; and that the male flowers really belong to these female 

 flowers may be shown by means of the highly characteristic pollen-grains 

 (Figs. 8 C, 9 C), which Renault l has discovered both in loco natali^ in the 

 loculaments of the anther, and in their place of destination, the apex of 

 the nucellus of the ovule, and with the same characteristic appearance in 

 both positions. Renault has figured three different male flowers, and 



FIG. 8. Male flower of a Cordaites, after Renault 2 . A transverse section. B longitudinal section of Cordaianthus 

 Penjoni, Ren., showing the elongated pollen-sacs disposed in tufts. C a single pollen-grain enclosing the internal 

 pluricellular cell-mass. 



described them as Cordaianthus Penjoni, C. subglomeratus, and C. Sapor- 

 tanus. These are the commonest species, the other and rarer kinds have 

 yet to be published. The flower-bud of C. Penjoni consists of a tolerably 

 thick axis closely enveloped in small narrow one-nerved spirally disposed 

 leaves. The summit of the axis is occupied by one or two male flowers, 

 and similar flowers, as is well shown in the transverse section 3 , are found 

 scattered about among the leaves, and appear in this case to be placed in 

 the leaf-axils. The individual flower, which is borne on a long cylindrical 



Renault 



a Renault (1). 



:1 Renault (1), t. 16, ff. 12, 15. 



