CAMPANULIN.E. 



563 



tnrued anteriorly (Fig. 602) (a position which is found to occur 

 within the Campanulacege). A twisting of the peduncle takes 

 place even before flowering (as in the Orchids) so that the ordin- 

 ary position of the 5-merous Dicotyledons appears to be restored. 

 The zygomorphy of the flower is especially present in the corolla, 

 which has a bipartite under-lip and a tripartite upper-lip, and is, 

 in Lobelia, anteriorly (apparently posteriorly) deeply cleft (Fig. 602). 

 There is 1 style, but the stigma is capitate and bilobed and sur- 

 rounded at its base by a whorl of hairs, which assists in pollination 

 (as a stylar-brush) in the same manner as the sweeping-hairs in the 

 Campanulacese and Composite. There is no terminal floiver in the 

 spicate, or racemose inflorescences. Lobelia has a capsule, several 

 others have berries. Isotoma (regular flower); Heterotoma has a spur; 



FIG. 602. Diagram of Lobelia 



fnlgens. FIGS. 603, 601. lobelia syphilitica. 



FIG. 603. Flower (Y). FIG. 601. Longitudinal section 

 of the same. 



Siplwcampylos ; Lysipoma (pyxidium) ; Clintonia (1-locular fruit). Metzleria 

 (all the petals are free). 



Entomophilous and protandrous. About 500 species, especially in the 

 Tropics; in this country, L. dortmanna (margin of lakes). Several are culti- 

 vated in gardens and conservatories as ornamental plants (Lobelia bicolor, 

 rimts,fulgens, etc., Siphocampylos, Centropogori). The latex of several species 

 of '1 upa is poisonous ; caoutchouc is also obtained from them. OFFICINAL : 

 "herba Lobelia" (the alkaloid lobeline) from the poisonous L. inftata, (N. 

 Am.). 



Order 4. Goodeniaceae. Chiefly Australian (200 species), closely related to 

 Orders 3 and 5, but without latex. The style is provided with a " collecting- 

 cup " which receives the pollen before the flower opens ; it has a small, hairy 

 aperture through which the pollen is forced out by the stigmas, and through 

 which they emerge when the pollen is shed ; it is sensitive and exhibits move- 



