442 MESSRS. P. GOTCH AND V. HORSLEY 



SECTION of both Posterior Columns and Lateral on opposite Side to Nerve. 



That is to say, this remaining effect represents 19/100 of the whole. This result 

 tallies with that obtained with the other method, which showed that the section of this 

 lateral only reduced the effect by about 20 per cent. This reduction in the cord 

 electrical change is thus a reduction in the number or intensity of the afferent impulses 

 which pass up the cord when the nerve is excited. Hence, as far as the method goes, 

 it would appear that the lateral column on the side of the stimulated nerve furnishes 

 a path, whether internuncial or not, we cannot say, but, at any rate, an redirect one 

 for the transmission, of 20 per cent, of the nerve impulses. 



B. Effect of Section of the Lateral Column on the opposite side to the Nerve Excited. 



The contradiction which exists between the results of the present research and the 

 interpretation given to their experiments by MIESCHER and WOROSCHILOFF is brought 

 into the strongest prominence by the consideration of the present group of results. If 

 the interpretation of these physiologists is warranted by their data, and is of general 

 application, then we should expect that the integrity or otherwise of the lateral column 

 opposite the entry of a stimulated nerve would have the preponderating influence upon 

 the passage of those nerve impulses which have entered the cord by the nerve, and 

 consequently upon the amount of the electrical change in the observed portion of cord. 

 We have, however, already accounted for 95 per cent, of the effect by showing that 

 the amount is dependent upon the integrity of the other columns, and this fact alone 

 is sufficient to show what a small share the integrity of the opposite lateral column 

 has in providing channels for the conduction of ascending impulses. 



We have made experiments upon five different animals, the results being separated 

 into those in which the lateral column was divided on the opposite side to the 

 nerve, and those in which all structures, except the opposite lateral column, were 

 cut 



