ADAPTATION. 235 



and excitation of an organ are carried on, must also be in 

 fluenced by this rhythm of action and reaction; and there 

 fore, after losing more than usual by the destructive process 

 they must gain more than usual by the constructive process. 

 But temporarily-increased efficiency in these appliances by 

 which blood and nervous force are brought to an organ, will 

 cause extra assimilation in the organ, beyond that required to 

 balance its extra expenditure. Regarding the functions as 

 constituting a moving equilibrium, we may say that diverg 

 ence of any function in the direction of increase, causes the 

 functions with which it is bound up to diverge in the same 

 direction; that these, again, cause the functions which they 

 arc bound up with, also to diverge in the same direction ; and 

 that these divergences of the connected functions allow the 

 specially-affected function to be carried further in this direc 

 tion than it could otherwise be further than the perturbing 

 force could carry it if it had a fixed basis. 



It must be admitted that this is but a vague explanation. 

 Among actions so involved as these, we can scarcely expect 

 to do more than dimly discern a harmony with first princi 

 ples. That the facts are to be interpreted in some such way, 

 may, however, be inferred from the circumstance that an 

 extra supply of blood continues for some time to be sent to 

 an organ that has been unusually exercised; and that when 

 unusual exercise is long continued a permanent increase of 

 vascularity results. 



69. Answers to the questions Why do these adaptive 

 modifications in an individual animal soon reach a limit? 

 and why, in the descendants of such animal, similarly condi 

 tioned, is this limit very slowly extended? are to be found 

 in the same direction as was the answer to the last question. 

 And here the connexion of cause and consequence is more 

 manifest. 



Since the function of any organ is dependent on the func 

 tions of the organs which supply it with materials and 



