GONOMYIA. 469 



vein, a little before or beyond it, the subcostal cross-vein placed at 

 its tip or near it ; the 2nd vein beginning about the middle of the 

 wing, arcuated, sometimes considerably so, or nearly straight, 

 sometimes simple, but generally widely though shortly forked.* 

 The 1st submarginal cell very short, more or less triangular, 

 owing to the anterior branch of the 2nd vein being so much 

 shorter than usual, and so obliquely placed as to appear almost 

 like a cross-vein, this appearance being more conspicuous in some 

 species than in others ; its petiole long, and its inner end generally 

 beyond the tip of the 1st longitudinal vein, or nearly opposite it ; 

 yet in the commonest Oriental species it is considerably anterior 

 to this tip ; marginal cross-vein absent ; the 3rd vein with a very 

 short or quite moderately long basal section, varying with the 

 species, remaining portion nearly or quite straight ; anterior cross- 

 vein varying in length, placed a little forward or backward, 

 generally in a line with the base of the 3rd vein and the proximal 

 side of the discal cell. Discal cell open or closed ; when open, it 

 coalesces with the 3rd posterior cell, which proves it is the 

 anterior branch of the 4th vein that is forked ; four posterior cells : 

 the 1st equal in length to, or shorter than, the 2nd submarginal 

 cell, varying in width with the species : the 2nd petiolate when 

 the discal cell is open, the petiole about as long as the cell ; when 

 the discal cell is closed, the 2nd and 3rd cells subequal, approxi- 

 mately normal in shape. Posterior cross-vein at some distance 

 before the proximal side of the discal cell, or in a line with it ; 

 the 5th, 6th, and 7th longitudinal veins nearly straight. 



Range. Europe,- Kirghis Desert, Central Africa, North America, 

 Brazil, the Orient, and Australia. 



In characterizing the genus Gonomyia,, Mg., in his ' Monograph 

 of Norih American TIPULID.E,' Osten Sacken described the genus 

 as possessing two submarginal cells, noting, however, two examples 

 in which the anterior branch of the 2nd longitudinal vein being 

 absent, the 1st submarginal cell was obliterated ; and he expressed 

 his opinion that if other species occurred with this character, a 

 new genus might be founded on them. He, however, retained 

 the form provisionally in Gonomyia, emending in a footnote 

 (p. 178) his definition of the genus to warrant the inclusion of 

 species with one submarginal cell only. 



In studying the question of the nomenclature of the veins in 

 the Gonomyia group, it will be seen that in this genus the 2nd 

 longitudinal veiu may (in the case of my two species G. incompleta 

 and G. flavomarginata) be considered to be forked as usual, but 

 the 3rd vein would, under this theory, be absent, and the anomaly 

 of the anterior cross-vein joining the 2nd and 4th veins, instead of 

 the 3rd and 4th as usual, would be seen. 



* In a North-American species (G. blanda, O. S.) it even forms a rectangle 

 at its base, with an appendix. 



2n2 



