CALYCULARIA RADICULOSA CAMPBELL 



45 



The genus Calycularia as generally understood comprises four species 

 of thallose liverworts of rather unusual distribution. One of these, C. laxa, 

 occurs in arctic Siberia, two, C. crispula and C. Birmensis, are found in India 

 and Burmah, while the other, C. radiculosa, occurs in Java. The question has 

 been raised whether the latter species really should be united with the 

 others. Schiffner 3 after an examination of the plant, concluded that it 

 should be placed in the genus Morkia, a genus sometimes regarded as a 

 section simply of the larger genus Pallavicinia. The material upon which 

 the present account is based was collected by the writer in Java in 1906 

 while staying at Tjibodas, one of the stations where the plant had originally 

 been collected. 



GENERAL MORPHOLOGY 



The plants are dioecious, the male plants ( Fig. 1 ) being decidedly smaller 

 than the females (Fig. 2). Antheridia may be developed while the plants 

 are not more than 5 mm. in length, but the male plants may reach a length 

 of 12-15 mm. The female plants are two or three times as long as the males 

 and may reach a length of about 30 mm., with a breadth of 'about 14 mm. 



Fig. 3. A, cross-section of the thallus, showing the thickened mid-rib and 

 rhizoids, r, x about 30. B, cells from the ventral side of the thallus, showing the 

 mycorrhiza, m, extending from the rhizoid, r, into the inner cells of the thallus, 

 x about 400. C, an inner cell of the thallus invaded by the mycorrhiza ; the nucleus 

 of the cell is still intact. D, oogonium-like enlargement of a mycorrhizal filament. 



3 Osterreichische Botanische Zeitschrift, Feb., 1901. 



