CALYCULARIA RADICULOSA CAMPBELL 61 



The spores are very different in their markings, as we have already 

 noted, and this seems to be .true also for Morkia. 



THE AFFINITIES OF CALYCULARIA RADICULOSA. 



Schiffner, from his study of Calycularia radiculosa concluded that it 

 should be removed from its present association with C. crispula, C. Birmensis 

 and C. laxa and united with Morkia. While there seems to be reason to 

 remove the species from the genus Calycularia, it may be questioned whether 

 its association with Morkia is justified. While Schiffner states that Morkia 

 is without conducting tissue in the mid-rib, Cavers has shown that in M . Flo- 

 towiana there are two strands of conducting tissue, but in Calycularia radicu- 

 losa these are entirely wanting. Moreover, the structure of the sporogonium, 

 i. e., the character of the thickenings of the wall and the markings of the 

 spores are quite different, and more like Makinoa, or some of the forms 

 usually referred to the Codoniacese. It would probably be better to con- 

 sider this plant as the type of a new genus intermediate in some respects 

 between Morkia and some of the less specialized forms like Makinoa or 

 Pellia. It is hardly likely that the line between the Codoniaceae and Lep- 

 tothceas (or Blyttiaceae, as Cavers has called them) is very well defined, 

 and it is probable that further study of the thallose Jungermanniales will 

 result in decided changes in the accepted arrangement of the genera. 



