86 WRIGHT : THE GEN US DIOSPYROS 



qocarpaj in I). Toposia the calyx of the male flower is closed 

 in the bud, and bursts irregularly into two or more segments 

 when the flower opens. The calyx segments are most 

 strongly marked in D. Embryopterii^ D., Ebenum, D. Gard- 

 neri, &c., but in many of the species belonging to this class 

 the conspicuousness of the calyx segments becomes reduced 

 during the development of the fruit. 



The calyces of the female and hermaphrodite flowers can 

 be divided into several classes, the first comprising those 

 which even after fruiting remain small, thin, and horizontal, 

 as in J). prjiriejis, D. montana, D. sylvatica, D. attenuata, D. 

 opcarjpa^and others, the second class comprising those which, 

 though they have undergone considerable enlargement 

 during the formation of the fruit, remain relatively thin 

 and horizontal, as in J). Moon ii^ and D. hirsuta, the third 

 those which increase considerably in size and form a shallow 

 or deep cup in which the ripe fruit is lodged, as in D. 

 Gardneri. D. oppositifolia, D. affinis, D. quaesita, D. insignis, 

 and many others. 



During the formation of the fruit the calyces of the third 

 class show different courses of development. The most 

 usual is to form a very large intersegmental pouch and 

 pointed api ces, together with a feeble reflex margin ; this is 

 best seen in D. quaesita; in other species, notably D. Ebenum, 

 the apices become strongly pointed and horizontal and the 

 margin strongly reflexed ; in D. insignis and D. crumenata 

 the enlarged woody calyx becomes strongly four-sided in 

 consequence of the dying back of the apices of the segments 

 and the very reflex nature of their margins. 



Many of the calyces, though highly pubescent in the 

 flower stage, become glabrous in the fruiting stage. The 

 hairs, with one exception, are long unicellular outgrowths 

 from single cells. In D. Ebenum, however, the base of the 

 shallow calyx cup is covered with unicellular and pedicel- 

 late hairs, the latter having very much the appearance of 

 globular multicellular glands supported on pedicels of 



