Neopycnocoma.] cxxii. EUPiiouBiACEiE (prain). 963 



ovule. It is, therefore, a matter for furtlier investigation as to whether the 

 conditions present do not indicate an abnormal condition of A, macrophylla, Pax, 

 showing pistillody, rather than the existence of a distinct genus. 



Ibk. KLAINEANTHUS, Pierre ex Prain in Kew Bulletin, 1912, 105. 



Flowers dicx3cious, apetalous. Male : Sepals 4-5, imbricate, connate 

 below in a campanulate tube as long as the lobes. Stamens 8 or 10, 

 2-seriate ; filaments of the outer series rather shorter than those of the 

 inner, surrounded by a ring of 4-5 fleshy ovate glands opposite the 

 sepals and alternate with the outer stamens ; anthers basifixed, smooth, 

 2-celled, dehiscence longitudinal, introrse. Rudimentary ovary 2-3-lobed, 

 as long as the stamens of the inner series. Female : Sepals 5, rarely 4, 

 imbricate, almost free to the base. Disk cupular with often an incom- 

 plete or complete ring of 5 or 4 staminodes inserted at the base of the 

 disk within. Ovary 3-celled, glabrous ; styles 8, glabrous, 2-partite to 

 their lower fourth, each arm obtusely 2-lobed at the tip. Fruit capsular, 

 3-coccous ; cocci crepitant, separating septicidally and also opening loculi- 

 cidally leaving a long trigonous persistent woody columella; epicarp 

 thinly fleshy ; endocarp thickly woody. Seeds large, flattened ovoid, 

 carunculate ; embryo rather large ; cotyledons broad, ovate-cordate, 

 flat. — A tree ; leaves alternate ; petioles pulvinate at the base and 

 apex ; flowers in ample terminal and axillary panicles ; males glomeru- 

 late, females subsolitary to a bract ; pedicels in both sexes slender, short, 

 articulate. 



A single endemic species. 



1. K. Gabonise, Pierre ex Prain in Kew BuUeiiii, 1912, 106. A 

 tree 30-00 ft. high, twigs reddish and pubescent when young, at length 

 grey and glabrous. Leaves distinctly petioled, oblong or obovate, bluntly 

 acuminate, margin entire, base wide- to narrow-cuneate, 4-8 in. long, 

 1J-3J in. wide, firm, dark green and dull above, paler and somewhat 

 polished beneath, glabrous on both surfaces; secondary nerves 8-12 

 pairs, raised beneath, looping within the margin ; petiole slightly pulvi- 

 nate at the base, distinctly pulvinate and geniculate at the apex, 1 J-2 in. 

 long. Panicles axillary and terminal, males as long as the females, 

 rather shorter than the leaves ; male flowers glomerate, females sub- 

 solitary ; pedicels in both sexes shorter than the calyx, distinctly articu- 

 late. Male: Calyx 5- or 4-lobed, companulate below; lobes oblong, 

 obtuse, pubescent and ciliate, strongly imbricate. Stamens 8 or 10, 

 2-seriate ; filaments of outer series shortest, surrounded by a ring of 4 

 or 5 extra-staminal glands alternate with the outer stamens and opposite 

 the calyx-lobes ; anthers basifixed, erect, introrse, dehiscence longitudinal. 

 Rudimentary ovary 3-gonous, as long as the inner stamens, 3- (rarely 2-) 

 lobed at the apex. Female : Calyx 5- (less often 4-) partite almost to 

 the base, otherwise as in the males. Disk cupular, denticulate and 4-5- 

 lobed. Staminodes often present in a complete or incomplete series, 

 hypogynous. Ovary 3-celled, glabrous ; styles 3, deeply 2-partite, the 

 arms again minutely and bluntly 2-fid. Capsule 3-celled, J in. across; 



