271] STUDIES ON GREGARINES WATSON 61 



spores rhombus-shaped (seen in one plane) and biconical, with but one 

 integument. 



Leger and Duboscq (1904:375) compared one of their new species 

 with the species in question. Their remarks are : 



"Stenophora chordeume nous parait, par sa forme, une espece tres voisine 

 de la Gregarine des Polydesmus et Fontaria des Etats-Unis, signalee par Crawley 

 (1903) sous le nom d' Amphoroides fontariae. Les figures qu'en donne cet auteur 

 dans sa PI. I fig. 12, 13, 14 nous portent a croire, d'apres les caracteres de 1'epi- 

 merite, qu'il s'agit plutot d'un Stenophora que d'un Amphoroides. II est d'ailleurs 

 impossible de se prononcer avec certitude sur ce point, car Crawley ne nous fait 

 pas connaitre les sporocystes de sa Gregarine, et on sait que, outre la forme de 

 1'epimerite, celle des sporocystes distingue nettement les Amphoroides des Steno- 

 phora; dans Amphoroides, ils sont biconiques; chez Stenophora, ils sont ovoides." 



Thus the basis for the original inclusion of the species in the genus 

 Amphoroides is not that of spore characteristics and until the spores are 

 known the generic position of the species will not be absolute. The shape 

 of the protomerite of the species under consideration is, however, very 

 unlike that of the type species of this genus, A. polydesmi Leger, and 

 hence the species cannot consistently be left in this genus. Its logical 

 position seems to be with the Stenophoridae because of elimination from 

 any other genus rather than from any positive character, and I should 

 designate it Stenophora fontaria (Crawley). 



STENOPHORA BROLEMANNI Leger and Duboscq 



[Figure 13] 



1903 Stenophora brolemanni Leger and Duboscq 1903a :339-40 



Stenophora : This gregarine is small, from 40 to 54//, long and is com- 

 pressed laterally, especially in the anterior part. It lives within the cell 

 of the host during the greater part of its life cycle. The older intercellu- 

 lar individuals are subspherical and occupy a cavity larger than that oc- 

 cupied by the younger ones, which is formed by the greater destruction 

 and compression of surrounding cells. The protomerite is invaginated 

 into the anterior end of the deutomerite like a cork into the neck of a bot- 

 tle. When the animal leaves the epithelium, the protomerite still retains 

 its invaginated position. The protomerite in profile is cylindrical, rather 

 flattened at the top, and when seen from the front it is as broad as high, 

 widest anterior to the middle and possesses at the summit a small plate 

 slightly concave upward and bearing in the center a small spherical pa- 

 pilla. Legr^r and Duboscq say this papilla may correspond to a protrac- 

 tile epimerite, for fibrillae seem to radiate from the apex outward over 

 the anterior third of the protomerite. The deutomerite seen in profile is 

 much larger at its posterior end than elsewhere, i. e., the animal is com- 



