326 THE FERTILISATION OF FLOWERS. [PART IIT. 
and rises up into the throat and so is accessible to the most short-— 
lipped insects. When the flower opens, the two divisions of the 
style are closely appressed and project into the lower part of the 
anther-cylinder which is filled with pollen (Fig. 112, 2). They are 
ee a a 
lt 
Fia. 112. 
1—5.—Achillea Millefolium, L. 
1.—A capitulum, from above. a, stigmas of female flowers; b, stigmas of florets of the disk iz 
their second stage; c, anther-cylinders of disk-florets in their first stage; d, bud about to open 
Numerous younger buds are hidden in the centre of the capitulum. : 
2.—Floret of the disk, just expanding; in section. e, ovary; f, ne x orice | the b 
of the style; g, style; h, tube of corolla; #, filaments; k, two divisions of the style ; 1, tip of st: yle 
Aan brush of hairs; m, anther-cylinder; m, throat of corolla; 0, valves, closing the anther 
cylinder. 
a disk-floret, slightly more advanced, with its bract; the pollen is issuing from the anth 
cylinder. 
4.—Older disk-floret. The stigmas are separate and exserted ; the anthers have been withdra 
into the corolla by contraction of the filaments. sot 7 
5.—Tip of style of a disk-floret in its first (male) stage. k’, stigmatic papille ; 1, sweeping-hair 
p, pollen-grains, 
6—8.— Chrysanthemum leucanthemum, L. 
6.—Disk-floret, in first (male) stage. 
7.—Ditto, in second (female) stage. 
8.—Tip of one stigmatic branch, seen from the inner side (x 60). 
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furnished at the tip with divergent hairs (5), so that as the sty 
elongates the pollen is pressed out at the upper end of the anther 
cylinder, whose terminal valves (0, 2) separate. Finally the 
branches of the style emerge, spread apart, turning their stigmat: 
SE > ht Rance NI has tiple Wing ac ‘oe 
