502 THE FERTILISATION OF FLOWERS. [PART III. 
alveolus, H ; (b) Sphinges : (23) Macroglossa fuciformis, L. (Stromberg), all 
sucking. See also No. 590, III. 
Ajuga pyramidalis, L., is said by Ricca to be homogamous 
(665); I have found it slightly proterandrous, and visited by 
humble-bees (609). 
Ajuga genevensis, L.—The flowers are larger than those of A. 
pyramidalis, and the nectary is very large. The plant is visited 
by humble-bees and by Lepidoptera. A proboscis 7 to 8 mm. long 
is requisite to reach the honey (609). 3 
REVIEW OF THE LABIATAE. 
Delpino mentions the following six points as characteristic of 
the flower in Labiatz (178). 
(1) The horizontal position of the axis of the flower. (2) 
Division of the corolla into upper and lower lips. (8) Union of 
two petals in the upper lip, three in the lower. (4) Position of 
anthers and stigma below the vaulted upper lip, which shelters 
them. (5) Position of the nectary at the base of the flower below 
the lower lip, whose anterior part forms a platform for insects. 
(6) Well-marked dichogamy (“ deciso asineronismo nello sviluppo 
degli organi sessuali”) (Delpino, No. 178, p. 128; Hildebrand, 
No. 360). 
But this summary is only correct if we call only those Labiates 
typical to which it directly applies; it requires most important — 
modifications if it is to hold good for the majority of Labiates. 
For (1) the axis of the flower is never or hardly ever truly 
horizontal, but, as a rule, is rather bent into the curve most 
convenient for long-tongued bees (Lamium, Galeobdolon, Gale- 
opsis, Betonica, etc.). (2) The upper lip is entirely absent in 
many cases (Ajuga, Teucrium), bracts belonging to the flowers 
above taking its place in sheltering the stamens from rain. (3) | 
The under lip is by no means always formed of three petals, but 
in Lamium, for instance, consists of one only; the two lateral 
petals have here assumed a special function, forming two erect 4 
lobes which receive the bee’s head between them. (4) The © 
stamens project beyond the corolla not only in Ajuga and 
Teucrium, where the upper lip is absent, but also in Thymus, 
Lycopus, Mentha, etc., and the stigma also projects beyond the 
a ee 
corolla in Salvia. (5) The fifth statement is correct for the great — 
