364 Menders Experiments 



fertilisation, since sixteen individuals are included in the 

 series. Therefore the participators in the fertilisation are 



Pollen cells AB-^AB^AB^AB^Ab-^Ab^Ab^Ab^ 

 aB-{-aB + aB + aB'hab-\-ab-\-ab + ab. 



Egg cells AB + AB-\-AB + AB + Ab^Ab + Ab-YAb + 

 aB + aB + aB^-aB + ab + ab-\-ab + ab. 



In the process of fertilisation each pollen form unites on an 

 average equally often with each ^gg cell form, so that each 

 of the four pollen cells AB unites once M^ith one of the 

 forms of ^gg cell ABy Ab, aB, ab. In precisely the same 

 way the rest of the pollen cells of the forms Ab, aB, ab 

 unite with all the other ^gg cells. We obtain therefore 



AB A^ AB^ AB^ Ab Ab Ab Ab 

 'AB^ Ab^ aB^ ~^'^ AB'^ Ab^ aB'^ ab^ 



aB aB aB aB ab ab ab ab 



AB Ab aB ab AB Ab aB ab' 

 or 



AB + ABb + AaB + AaBb + ABb ■\- Ab -V AaBb + Aab + 

 AaB + AaBb + aB + aBb + AaBb + Aab + aBb -\-ab = AB + 

 Ab + aB^-ab-{-2ABb + 2aBb + 2AaB-{-2Aab + ^AaBb'^, 



In precisely similar fashion is the developmental series 

 of hybrids exhibited when three kinds of differentiating 

 characters are conjoined in them. The hybrids form eight 

 various kinds of ^gg and pollen cells — ABC, ABc, AbC, 

 Abe, aSC, aBc, abC, abc — and each pollen form unites 

 itself again on the average once with each form of ^gg cell. 



The law of combination of different characters which 

 governs the development of the hybrids finds therefore its 

 foundation and explanation in the principle enunciated, 

 that the hybrids produce ^gg ceils and pollen cells which 

 in equal numbers represent all constant forms which result 

 from the combinations of the characters brought together 

 in fertilisation. 



* [In the original the sign of equaHty (=) is here represented by 

 evidently a misprint.] 



