Increasing Quantities of Fat in Diet on Metabolism in Dogs. 271 



(c) Another period of five days was next investigated, when the diet 

 was increased still more to 51-73 grams of fat. The quantity of urine 

 now passed varied from 96 to 130 c.c., with a specific gravity of 1040 

 to 1052. The nitrogen was only analysed in the urine during three 

 days out of the five in consequence of an unfortunate accident. It 

 varied from 4*307 to 4*971 grams, and remained pretty constant during 

 this period. The dog daily passed his faeces, and the quantity in con- 

 sequence appears smaller than in the preceding cases, varying from 

 19*69 to 54*38 grams; at the same time the quantity of nitrogen con- 

 tained in the faeces varied from 0*322 to 0*918 gram. On the third 

 day of this period the fat in the faeces was lost; on the other days the 

 quantity varied from 3*050 to 1*602 grams. 



With this we get a varied absorption of proteids, the quantity vary 

 ing from 84*83 to 94*68, while as far as the fat is concerned it varied 

 from 94-10 to 98'93 per cent. 



Having considered this table in detail, we can now consider the 

 average of the three periods (), (b), (c) in this case, where partial removal 

 of the large intestine had been carried out. 



It was found in this partial removal, even after shrinkage of the part 

 isolated, that over a half of the total length of the large intestine had 

 been removed. The dilatation of the rectum accounts for the retention 

 of fasces on some days so as to cause constipation. 



From Table V the average of the three periods, the addition of an 

 increasing quantity of fat being added to a fixed nitrogen diet, is seen 

 to cause a decrease in the quantity of urine, the amount falling from 

 172 to 169 c.c., and then to 112 c.c. The specific gravity did not 

 quite coincide, as it did not rise steadily in the three periods. 



As far as the nitrogen in the urine is concerned, we see also that as 

 the fat was increased, so the quantity of nitrogen in the urine fell 

 from 5-596 to 4*991 grams and 4-680 grams. So that as in the normal 

 dog the nitrogen sparing properties of the fat are well brought out. 



The quantity of the faeces for all practical purposes is not much 

 influenced by increasing the quantity of fat, and certainly not in the 

 degree which would seem to occur in the two normal dogs, in both of 

 ch, on the fat being increased in the diet, the quantity of faeces 

 'ere augmented. The variation may be in part explained by the con- 

 stipation which occurred in periods (a) and (b). 



The nitrogen of the faeces in this dog, if anything, was decreased in 

 quantity by increasing the fat, for whereas during the first five day?, 

 period (), 0*792 gram of nitrogen was daily eliminated in the faeces, 

 on the diet being increased to 36 -7 3 grams of fat during the four days 

 of period (b), the nitrogen was increased to 0*614 gram, and during 

 period (c), when no less than 51-73 grams of fat were being taken, the 

 nitrogen amounted to 0*624 gram. 



With regard to the fat in the faeces, the table shows that the quan- 



VOL. LXIV. Y 



