Increasing Quantities of Fat in Diet on Metabolism in Doys. 281 



The quantity of faeces during the two periods is 83'78 and 83'49 

 grams, so that the increase of fat has undoubtedly caused in this dog 

 an increased quantity of faeces. The nitrogen during these two periods 

 was 1'098 and T088 grams, so that the increase in the quantity of 

 fat in the diet has caused a slight increase in the quantity of nitrogen 

 in the faeces, the same as we find in the normal dogs. The quantity 

 of fat in the two periods was 0-769 and 0'873 gram, that is to say, a 

 slight tendency to an increase, through the increased quantity of fat in 

 the diet. 



We thus see that the dog after total removal of the large intestine 

 shows a marked increase in the quantity of nitrogen in the faeces ; 

 at the same time, in spite of this increase, it can be still more 

 increased by increasing the fat given in the diet. As far as the 

 fat is concerned the quantity of fat in the normal dog's fseces, as com- 

 pared with that after removal "of the large intestine, is practically the 

 same. It is thus well borne out that as far as the fats are concerned 

 they are well absorbed by the small intestine, the large intestine having 

 no influence whatever on their absorption. 



On the other hand, as far as the proteids are concerned, they are not 

 so well absorbed at least 10 per cent, less but there is an increase 

 in the quantity of proteid or nitrogen in the faeces of the dogs in which 

 the large intestine has been removed, and that this increase is slightly 

 augmented by increasing the quantity of the fat in the diet, as in 

 normal dogs; at the same time the decrease in absorption, as thus 

 indicated in normal dogs, is not so marked, if at all marked, after 

 removal of the large intestine. 



Having considered the general metabolism, and having seen how the 

 quantity of faeces is increased in dogs in which the large intestine has 

 been removed, we now come to consider the faeces more carefully as 

 regards their general appearance and the quantity of water. 



The Influence of Diet on the Daily Quantity and the Amount of Water in the 

 Fceces when the Large Intestine is Removed. 



Immediately after the operation, the dogs for the first few days took 

 practically no food, and then were put on a milk diet, which conse- 

 quently caused the light motions so typical of this diet. When the 

 diet was slowly converted into that of biscuit and meat, on which a 

 normal dog would be passing well-formed faeces, the dog continued 

 to pass a semi-fluid motion. In the case of the partial removal of the 

 large intestine this was well marked, so that the dog for the first few 

 weeks suffered from more or less diarrhoea. Before death, however, 

 throughout the time in which metabolism experiments were being 

 done, the dog passed faeces almost of normal consistence, and, as is 

 seen by the post mortem, this is ^to be explained by the sack-like 

 enlargement of the rectum. 



