Increasing Quantities of Fat in Diet on Metabolism in Dogs. 305 



In normal dogs it is found that the increase of fat in the diet causes 

 a progressive increase in the total quantity of water eliminated in the 

 faeces, while the percentage of water is decreased, the total quantity 

 of the faeces increasing with the increase of fat in the diet. 



In partial removal of the large intestine there is also an increase in 

 the total quantity of water, as well as apparently an increase in the 

 percentage of water. 



In complete removal of the large intestine there is an increase in 

 the total quantity of water eliminated in the faeces, with an increase 

 in the percentage of water eliminated by the bowel, the faeces increasing 

 in quantity with the augmentation in the quantity of fat in the 

 diet. 



It is thus seen that increase of fat causes an increased quantity of 

 faeces in the normal dog, the increased quantity of faeces is accompanied 

 by an increase of total water, but a decrease in the percentage of 

 water eliminated in the faeces, while in the case of the removal of the 

 large intestine, the increase in the quantity of faeces is accompanied 

 both with an increase in the total quantity of water and the percentage 

 of water. (Table XIV.) 



6. The influence of the removal of the large intestine on the absorp- 

 tion from the alimentary canal. 



It has been already seen that the dogs passed a larger quantity of 

 faeces when the large intestine had been removed, and this is to a small 

 extent brought out in partial removal of the large intestine. It is 

 seen that the increase in quantity of faeces is principally due to the 

 increase in quantity of water, the total quantity being nearly five 

 times as much as in the case of the normal dogs. The quantity of 

 nitrogen in the faeces is increased to nearly three times as much as in 

 the normal dogs, while the quantity of fat remains unaltered, and in 

 those cases which were examined the faeces contained no carbo- 

 hydrates. 



It is therefore seen that as far as the absorption from the alimentary 

 canal is concerned : 



(a) The carbohydrates are absorbed equally well with and without 

 the presence of the large intestine. 



(b) The fats are also absorbed equally well. The normal dogs show 

 a percentage of absorption from 94 to 98, according to the amount of 

 fat given. This apparently better absorption occurs with the increase 

 of fat in the diet. 



In partial removal of the large intestine the percentage is roughly 

 96, while when the entire large intestine is removed, from 92 to 98 

 per cent, of the fat given is absorbed, so that the dogs with and without 

 the large intestine appear to absorb fat equally well. 



(c) The proteids, as indicated by the nitrogen in the faeces, are, how- 

 ever, very markedly influenced. In normal dogs 93 to 98 per cent. 



