THE PROBLEM OF INBREEDING 113 



In the same way, 



= 100 (4-2) m 50 _ 



Z 2 = l°°f^ - 75. 



= 100116-2) _ g7 s 

 16 



These results may be expressed verbally in the 

 following way : In the last two ancestral genera- 

 tions x is 50 per cent inbred ; in the last three 

 generations it is 75 per cent inbred ; and in 

 the last four generations it is 87.5 per cent 

 inbred. 



This pedigree table and the constants will 

 repay further consideration, since the case is a 

 limiting one. With the table at hand it is possible 

 to grasp a little more clearly the precise meaning 

 of the coefficients of inbreeding. Thus it is seen 

 that what the value of Z x = 50 really signifies 

 is that because the individuals a and b were 

 brother and sister the number of different ancestors 

 which x can possibly have in any ancestral genera- 

 tion cannot be more than 50 per cent of the total 

 number theoretically possible for the generation. 

 That is, x's sire and dam having been brother and 

 sister means that x cannot have had more than 

 2048 different great-great-great-great-great-great- 



