NAMED RAn-LESlA. 383 



Eupomatla} which I have placed near that natural 

 order. 



Another point to be inquired into connected with the 

 same subject is, in wliat manner the impregnation of the 

 female flower is likely to be effected by anthera} so com- 

 pletely concealed as those of Bafflesia seem to be in all 

 states of the flower ; for it does not appear either that they 

 can ever become exposed by a change in the direction of 

 the limb under which they are inserted, or even that this 

 part of the column in any stage projects beyond the tube 

 of the perianthium. 



It is probable, therefore, that the assistance of insects is 

 absolutely necessary ; and it is not unlikel}^ both as con- 

 nected with that mode of impregnation and from the 

 structure of the anthera itself, that in Bafflesia the same 

 economy obtains as in the stamina of certain Aroidccc, in 

 which it has been observed that a continued secretion and [~^i7 

 discharge of pollen takes place from the same cell ; the 

 whole quantity produced greatly exceeding the size of the 

 secreting organ. 



The passage of the pollen to the bottom of the flower, 

 where it is more easily accessible to insects, seems likewise 

 to be provided for, not only by the direction of the anthera?, 

 but also by the form of the corresponding cavities in the 

 neck of the column, in the upper part of Avhich they are 

 immersed. 



That insects are really necessary to the impregnation of 

 Bafflesia, is confirmed by Dr. Arnold's statement respecting 

 the odour of the plant, by which they may be 3U})posed to 

 be attracted, and also by the fact of the swarms actually 

 seen hovering about and settling in the expanded flower. 



The structure of Bafflesia is at present too imperfectly 

 known to enable us to determine its place in the natural 

 system. I shall, however, offer some observations on this 

 question, which can hardly be dismissed without exami- 

 nation. 



As to which of the two primary divisions of phaenoga- 

 * riinders's Voyage^ ii,^. 597 {Ante^p. 73), tub. 2. 



