496 ON THE ORGANS AND MODE OF FECUNDATION 



year, tliat he considers the analogous glands, existing in 

 most other tribes of Orchidese, as equally belonging to the 

 stamen : in his " Introduction," however, he refers them 

 to the stigma in all cases exce})t in Ophrydea?. 



ToAvards the end of 1830 the first part of Mr. Francis 

 Bauer's lilustrations of Orchideous Plants edited by Mr. 

 Tin (Hey, was published. 



From this work, of the importance and beauty of which 

 it is impossible to speak too highly, it may be collected 

 that Mr. Bauer's opinion or theory of impregnation in Or- 

 chicleso does not materially differ from that of Batsch, 

 Richard, and other more recent writers. From one of the 

 figures it appears that tliis theory had occurred to him as 

 early as 1792; and in another figure, bearing the same 

 date, he has accurately represented the structure of the 

 grains of pollen in a plant belonging to Ophrydese, a struc- 

 ture which I had not ascertained in that tribe till 1806. 

 Althouo;!! Mr. Bauer's theory is essentially the same as 

 that of Batsch and Richard, yet there are some points in 

 which it may be considered peculiar ; and chiefly in his 

 supposing impregnation to take effect long before the ex- 

 cos: pansion of the flower, at a time when the sexual organs 

 are so placed with relation to each other that the fecun- 

 dating matter, believed by him to pass from the pollen 

 mass through its caudicula, where that part exists, to the 

 gland attached to it, may be readily communicated to the 

 stigma, v>dtli which the gland is then either in absolute con- 

 tact or closely approximated. The more important points 

 of this account may be extended to nearly the whole order, 

 but is strictly applicable only to Satyrina3 or Ophrydeae, a 

 tribe in which Mr. Bauer seems, with Mr. Lindley, to con- 

 sider the glands as belonging to the stamen and not to the 

 stigma.^ In those genera of this tribe in which the glands 



1 In the second part of Mr. Bauer's Illustrations, which lias appeared since 

 this paper was read, the explanation of Tab. 3, fig. 6, is corrected in the follow- 

 ing manner : 



" For 6. A pollen mass with its caudicula and gland taken out of the anther; 



"HeaclG. A pollen mass with its caudicula and the internal socket of the 

 stigmatic gland." 



It is evident, indeed, in the second part of the Illustrations, from figs. 8, 9, 

 11, and 12, of Tab. 12, representing details of ^^atyrlvm puatidatim, and the 



