18 ON THE PROTEACEiK OF JUSSIKU. 



concave tops of the laciniae of the calyx ; all considerable 

 deviations from which may safely be employed in charac- 

 30] tensing genera. In this way Bhopala, Xylomelum, 

 and Lamberila are readily distinguished from Embothfium, 

 GrewHea, and Hakea ; and thus also Persoonia and Bra- 

 bejum remarkably differ from Guevina ; while Bellendena 

 differs from all others in having its stamina distinct from 

 the calyx, affording however an indication of the real origin 

 of these organs in the whole family. 



The deviations from the usual structure of anthers in 

 this order are not many ; but some of them are of so sin- 

 gular a nature as to constitute the essential characters of the 

 genera in which they take place. These genera are Simsia, 

 Conospermum, and Synaphea, all of which are most truly 

 syngenesious ; for not only do their antherae firmly cohere 

 together, but the corresponding lobes of these being, when 

 considered separately, entirely open, are so applied to each 

 other as to form but one cell, without a trace of any in- 

 termediate membrane. In Simsia the four antherae are 

 perfect, each consisting, as in the rest of the order, of two 

 lobes, and therefore the whole before bursting constitute 

 four cells. Whereas in Conospermum and Synaphea one 

 filament is entirely barren, the two lateral ones have each a 

 single-lobed anthera, and the fourth alone is perfect : hence 

 before bursting the whole form only two cells. 



This remarkable structure, which can only be ascer- 

 tained before the opening of the calyx, necessarily escaped 

 Dr. Smith in describing his Conospermum, for I conclude 

 he had only the expanded flower before him, and the 

 appearance of the antherae in this state after their separa- 

 tion justifies him in referring the genus to Tetrandria : but 

 according to the view now given of its structure, it can 

 have no other pretension to a place in this class than its 

 belonging to Proteaceae ; and the order Syngenesia Mono- 

 gamia being abolished, it must be referred to Triandria. 



The only remaining anomaly in these parts occurs in 

 si] Franklandia, and consists in the anthera, or rather that 

 portion of the filament on which it is fixed, adhering to 

 the calyx through its whole length. 



