76 MORPHOLOGY OF PAPILIONACEOUS FLOWERS. [BOOK i. 



tubercles on the pistillary border,, which subsequently form 

 ten summits of distinct styles, and which correspond in pairs 

 to five ovaries, the centre of which also becomes hollowed by 

 a fissure, which forms its cavity without any change being 

 externally apparent. 



The necessary conclusion from all these observations is, 

 that the parts, from their earliest appearance, present the 

 relations of adhesion which they subsequently exhibit in the 

 perfect flower. The monophyllous calyx on its first appear- 

 ance was a body simple at the base. The petals coherent by 

 their base with the staminal tube, originated from a base 

 common to them with the stamens, and the latter at their 

 origin were united by this base in the same manner as they 

 appear subsequently. The ovaries were from the first grouped 

 and adherent together, nearly in the same manner as the 

 flower subsequently exhibits them, their styles being distinct 

 at the summit, coherent in the rest of their extent, which has 

 been more slowly developed. As regards the peculiar results 

 to be deduced from these observations relative to the sym- 

 metry of the flower of the Malvaceas, we have noticed them 

 above, and it would be useless to repeat them. (Annals of 

 Natural History, xvi.) 



B. The progressive development of the Papilionaceous Flower, 

 among Leguminous Plants, by Schleiden and Vogel : 



1. The flowers are at their origin perfectly regular. 



2. The subsequently cohering parts originate as free points, 

 are developed free, and cohere subsequently. 



3. All the parts of the flower are at their first appearance 

 green leaves. 



4. Even in the earliest stage only one carpellary leaf is visible 

 in the Leguminosse, which is open in the direction of the axis. 



5. The anthers are formed from leaves, the inner cellular 

 tissue being converted in part into pollen; and the loculi 

 originate at both sides of the margin of the leaf, which is 

 subsequently changed into the bursting rima. 



6. The ovules are formed alternately at the upper margin 

 of the ovarium, and consist of the nucleus and generally of 

 two integuments, rarely of an integumentum simplex. 



