Genus of T'lpuWdsd from Turkestan, 581 



D. The Indian Oonomyia incompleta, Biun., and flavo- 

 tnarginata^ Brun., and the African Atarha lamellar is, Speis., 

 belong to the genus Ltponeura, Skuse, which is distinct 

 from Atarba, O.-S. Of the five American species placed by- 

 Alexander (Ent. News, 1912, p. -41 S) in Liponeura, which 

 he considers a subgenus of Gonomyiu, four doubtless belong 

 there; but, although I have not seen the " more extensive 

 discussion of the genus published elsewhere," of which 

 Alexander speaks, 1 venture to express some doubt whether 

 Oonomyia nianca, O.-S., really is congeneric with the others. 

 Osten-iSacken says in his description: ** the venation is 

 precisely like that of G. sulphur ell a, except that the posterior 

 branch of the second longitudinal vein is obliterated ; thus 

 the second longitudinal vein, shortly before its tip, takes 

 a sudden turn towards the anterior margin, in consequence 

 of which the submarginal cell is trumpet-shajied y This 

 description is evidently correct, although Osten-Sacken later 

 (Berl. ent. Z. 1887, p. 165 and 202), by some mistake, stated 

 that the anterior branch of the second vein is obliterated in 

 G. manca — a statement in which he was followed by Brunetti. 

 It was clearly owing to this sudden turn of the second vein 

 that Osten-Sacken could recognize a Gonomyia in his insect, 

 and any Gonomyia with E3 (the posterior branch of the 

 " second " vein) obliterated could be recognized by the fact 

 that Ro has retained its original shape and position with the 

 apex suddenly upturned, forming an angle with the basal 

 part of R2 and ending a little in front of the apex of Rj. In 

 Liponeura the vein R2+3 is almost perfectly straight or gently 

 and slightly curved, and the apical part does not form an 

 angle with the basal part, and ends somewhat farther from 

 the tip of Ri, thus occupying a position intermediate between 

 R2 ai'd R3 in the typical Gonomyics. The difference may 

 seem to be of little importance ; but it indicates that the 

 disappearance of the first submarginal cell of the old nomen- 

 clature in Gonomyia manca is due to a simple obliteration 

 of R3, whereas in Liponeicra it is due to an actual fusion of 

 R2 and R3, of the same kind that has taken place in all 

 Limnobiina and Rhamphidiina. I therefore think that 

 Gonomyia vianca is a true Oojiomyia, for which not even a 

 subgenus is necessary ; whereas Liponeura should hold the 

 position assigned to it by Skuse as a distinct genus of the 

 Rhamphidiina near yl^ar^a, O.-S., the latter genus now being 

 restricted to its type, A. picticornis, O.-S., and the Indian 

 A. flava, Brun., which is clearly congeneric. This opinion 

 is strengthened by the different structure of the male pro- 



