xxx] 



CODONOTHECA 



125 



are seen in fig. 427, 2, 3, on the inner face of the lobes. Some of 

 the specimens have a fairly long pedicel : in the example shown 

 in fig. 427, 1, the fleshy part of the basal portion is not preserved, 

 only the more resistant vascular core. In a later account of these 

 organs Sellards speaks of several lying by the side of a central 

 stalk to which he thinks they were originally attached by slender 

 pedicels. In view of Dr Benson's interpretation of the morpho- 

 logy of Telangium it is permissible to suggest that if a central 



FIG. 427. Codonotheca caduca. 1. The vascular tissue preserved as a conical 

 base which was originally surrounded by fleshy tissue (cf. 5); I III, VI, 

 vascular bundles (x 2). 2, 3. Segments with spores; c, floor of cup (nat. 

 size). 4. Vascular tissue as a cone in the middle of the fleshy base (nat. 

 size). 5. Plan of the whole organ opened out flat (nat. size). 6. Spores 

 (x 28). 8. A single spore showing median slit (x 84). (After Sellards.) 



sporangium in such a synangium as that of Codonotheca developed 

 a megaspore and the peripheral sporogenous lobes were sterilised, 

 the result would be an arrangement not unlike the apical region 

 of the seed Physostoma, the tentacles of which have been homo- 

 logised with the canopy of Lagenostoma. There are obvious diffi- 

 culties in the way of this, perhaps strained, comparison: the 

 larger size of the spore-bearing linear segments of Codonotheca 



