XXXVIIl] BEANIA 503 



borne in pairs on the adaxial side of the terminally expanded 

 pedicels are covered with small granulations which Nathorst 

 thinks may be clusters of microspores, the apparent seeds being 

 'antherangia.' The granulations are, however, very similar to 

 those on the larger detached seed-like bodies originally described 

 by Nathorst from Rhaetic beds in Sweden as Antherangiopsis 

 rediviva 1 : subsequent examination of that species demonstrated 

 that the granulations are due to the presence of resinous bodies 

 in the tissues of true seeds 2 , and it is not improbable that a similar 

 interpretation may hold for the surface-features in the supposed 

 male organs of Beania Carruthersi. Pending further evidence it 

 may be suggested that Beania Carruthersi is like B. gracilis a 

 seed-bearing shoot. The Rhaetic specimens described by Nathorst 

 as Stenorrachis scanicus 3 are similar in habit to Beania but differ 

 in the forking of the sporophylls (fig. 656) and in the absence 

 of any terminal swelling on which the seeds are borne : Nathorst 

 considers that Stenorrachis may be the female organ of a Nilssonia 

 and it is not improbable that that genus and Beania are closely 

 allied types. We have no definite information with regard to 

 the reproductive organs of the Nilssoniales : the closer resem- 

 blance which their fronds bear in the structure of the epidermal 

 cells to those of recent Cycads is consistent with the view that 

 their fertile shoots were also more like those of existing types. 

 It is, however, still an unsettled point whether Beania is more 

 closely allied to the Cycadophyta or to the Ginkgoales, but the 

 balance of opinion is in favour of the former alliance. 



Zamiostrobus. Endlicher. 

 Cycadeostrobus. Carruthers. 



Though instituted by Endlicher 4 for a cone figured by Lindley 

 and Hutton as Zamia macrocephala 5 which is almost certainly 

 Abietineous and has no claim to be included in the Cycadales, 

 the genus Zamiostrobus has been adopted by many authors for 

 Cycadean ovulate cones, not only such as are believed to be 

 closely allied to those of Zamia but for Cycadean cones generally. 



1 Nathorst (02) p. 20, PI. i. figs. 22, 23. 



2 Ibid. (09 2 ) p. 23. 3 See Vol. iv. 



4 Endlicher (40) p. 72. See also Schimper (72) A. p. 201. 



5 Lindley and Hutton (35) A. PL 125. 



