EAR AS AN ORGAN FOR SOUND SENSATIONS. 



367 



the tympanic membrane are communicated to the tip of the handle 

 of the manubrium. As the handle moves in, the chain of bones 

 makes a rotary movement around an axis which may be defined as 

 the line passing through the attachment of the short process of the 

 incus and the anterior ligament of the malleus. The general posi- 

 tion of this axis is represented by the line a-b in Fig. 164. This line 

 passes through the neck of the malleus ; so that as the handle moves 

 in the head of the malleus and the upper part of the incus move in 

 the opposite direction, while the long process of the incus together 

 with the stapes, being below the axis, move in the same direction aa 

 the handle, (see Fig. 164A). The chain of bones, therefore, acts 

 like a bent lever whose fulcrum is at a, the power arm being repre- 



Fig. 164. To illustrate the lever 

 action of the ear bones (McKendrick) : 

 M, The malleus; e, the incus; a-b, the 

 axis of rotation: a, short process of 

 incus abutting against the tympanic 

 wall ; a-p, the power arm ; a-r, the 

 load arm of the lever. 



Fig. 164 A. Schema to illustrate the 

 way in which the ear ossicles act to- 

 gether as a bent lever in transmitting the 

 movements of the tympanic membrane 

 to the membrane of the fenestra ovalis. 

 1, The handle of the malleus; 2, the long 

 process of the incus; 3, the stapes ; a-b, 

 the axis of rotation. 'Ihe arrows indi- 

 cate a movement inward of the tympanic 

 membrane. 



sen ted by the line p-a and the load arm by the line r-a. According 

 to Helmholtz,* the distance p-a is equal to 9.5 mms., while r-a 

 is 6.3 mms. The movement at r, therefore, or the movement 

 of the stapes, will have only two-thirds of the amplitude of the 

 movement at p, but will have a correspondingly greater force 

 (one and one-half times). The mechanisms of the tympanic 

 membrane and the ear bones combine, therefore, to convert the 

 vibratory movements of the tympanic membrane into smaller 

 but more intense movements of the membrane of the fenestra 

 ovalis. It should be borne in mind, however, that the amplitude 



* Helmholtz, "Die Lehre von den Tonempfindungen, etc.," fifth edition, 

 1896. See also English translation by Ellis. 



