54 PROTOMYXA AND THE MYCETOZOA LESS. 



and throughout the substance of which are found numerous 

 nuclei. In this condition they creep about over bark or 

 some other substance : and in doing so ingest solid food 

 (Fig. 7, A). It has been proved that they digest protoplasm : 

 and in one genus pepsin the constituent of our own gastric 

 juice by which the digestion of proteids is effected (see p. 12) 

 has been found. They can also digest starch which has been 

 swollen by a moderate heat as in our own bread and rice- 

 puddings but are unable to make use of raw starch. 



After living in this free condition, like a gigantic terrestrial 

 Amoeba, for a longer or shorter time, either a part or the 

 whole of the protoplasm becomes encysted l and breaks up 

 into spores. These (B) consist of a globular mass of proto- 

 plasm covered with a wall of cellulose : the cysts are also 

 formed of cellulose. 



By the rupture of the cell-wall of the spore (c) the proto- 

 plasm is liberated as a flagellula (D) provided with a nucleus 

 and a contractile vacuole, and frequently exhibiting amoeboid 

 as well as ciliary movements. After a time the flagellulae 

 lose their cilia and pass into the condition of amoebulae (E), 

 which finally fuse to form the plasmodium with which 

 we started (F H). In the young plasmodia (c 1 ) the 

 nuclei of the constituent amcebulae are clearly visible, and 

 from them the nuclei of the fully developed plasmodia are 

 probably derived. It would seem, therefore, that in the 

 fusion of amoebulae to form the plasmodium of Mycetozoa the 

 cell-bodies (protoplasm) alone coalesce, not the nuclei. 



There is a suggestive analogy between this process of 



1 The process of formation of the cyst or sporangium is a compli- 

 cated one, and will not be described here. See De Bary, Fungi, 

 Mycetozoa, and Bacteria (Oxford, 1887), and Lister, Catalogue of the 

 Mycetozoa (London, 1894). 



