CHAP. I.] 'SYMPATHETIC'-SUBMAXILLARY SALIVA. 31 



found by Eckhard to be 1004'6, that flowing on stimulation of the 

 sympathetic had a specific gravity of 101 5'6. The amount of solid 

 matter in the sympathetic isa'liva of the dog was found by Eckhard 1 

 to be 2'7 per cent., by Heidenhain sometimes to be as much as 

 3'74 and 5'86 per cent. On long-continued stimulation, Heidenhain 

 found that the solid matters decreased so that the saliva 2 acquired 

 the characters of that secreted under the influence of stimulation 

 of the chorda tympani. This is in accordance with the facts given 

 above with regard to the gradual exhaustion of the mesostate of the 

 gland-cells. 



Differences Langley has shewn that the effects of stimulating the 



in effect of chorda tympani and sympathetic of the cat are different 

 stimulating from those observed in the case of the dog. In the cat, 

 nerves in Cat sympathetic saliva is in most cases less viscid than the 

 and Dog. chorda saliva. Moreover whilst atropin paralyses the chorda 



secretion in the dog, leaving the sympathetic secretion unaltered, in the 

 cat it paralyses the sympathetic secretion as well as that of the chorda 3 . 

 From this it would appear that relatively more secretory fibres run by 

 the sympathetic in the cat than in the dog. 



The following analyses shew the usual difference in percentage com- 

 position between chorda- and sympathetic-saliva in the cat (Langley 4 ). 



I. 



(1) Saliva obtained by weak stimula- ) 



Percentage PprppTltaffp Total 

 of organic ercen g e p ercentage 

 substances. ' r asn ' of solids. 



n , -I -I n, ,v ,. 



tion ot the lett sympathetic 



(2) 5 mgrm. atropin given; saliva ob- 

 tained by strong stimulation of } 0-525 0-454 0-979 

 right sympathetic 



iva ob- | 

 :ion of > 



II. 



(1) Saliva obtained by stimulating | 



chorda, shocks fairly felt by tip V 0-865 0-339 1-205 

 of tongue 



(2) Saliva obtained by somewhat 



stronger stimulation of sympa- J- 0-426 0-275 0-701 

 thetic 



1 Eckhard, ' Ueber die Unterschiede des Trigeminus- und Sympathicusspeichels der 

 Unterkieferdriise des Hundes.' Beitrcige zur Anat. u. Phys., Vol. n. p. 205. 



- Heidenhain, * Physiologic der Absonderungsvorgange.' Hermann's Handbuch, 

 Vol. v. Part i. p. 48. 



3 Langley, ' On the Physiology of the Salivary Secretion. Part i. The influence of 

 the Chorda Tympani and Sympathetic Nerves upon the secretion of the sub-maxillary 

 gland of the Cat.' Journal of Physiology, Vol. i. (1878), p. 96 et seq. 



4 Langley, 'On the Physiology of the Salivary Secretion.' Part m. Journal of 

 Physiology, Vol. vi. p. 92. 



