428 LIFE OF PROFESSOR HUXLEY 



nerves, may represent a pair of prae-oral visceral arches. 

 In his unpublished notes there is evidence that he was 

 bringing to the support of this conclusion the discovery 

 of a supposed 4th branch to the trigeminal nerve — the 

 relationships of this (which he proposed to term the 

 " hyporhinal " or palato-nasal division) and the ophthalmic 

 (to have been termed the " orbitonasal " i) to the trabecular 

 arch and a supposed prae-mandibular visceral cleft, being 

 regarded as repetitional of those of the maxillary and 

 mandibular divisions to the mandibular cleft. So far a& I 

 am aware, von Kupfter is the only observer who has given 

 this startling conclusion support, in his famous Studien 

 (Hf. I. Kopf Acipenser, Miinchen, 1893), and from the 

 nature of other recent work on the genesis of parts of the 

 cranium hitherto thought to be wholly trabecular in origin, 

 it might well be further upheld. As for the discovery of 

 the nerve, I have been lately much interested to find 

 that Mr. E. Phelps Allis, jun., an investigator who has 

 done grand work in Cranial Morjihology, has recently 

 and independently arrived at a similar result. It was 

 while working in my laboratory in July last that he 

 mentioned the fact to me. Remembering that your 

 father had published the aforementioned hints on the 

 subject, and recalling conversations I had with him, it 

 occurred to me to look into his unpublished MSS. (then 

 being sorted), if perchance he had gone further. And, 

 behold ! there is a lengthy attempt to write the matter 

 up in full, in which, among other things, he was seeking 

 to show that, on this basis, the mode of termination of 

 the notochord in the Craniata, and in the Branchiostomidai 

 (in which the trabecular arch is undifferentiated), is readily 

 explained. Mr. AUis's studies are now progressing, and 

 I have arranged with him that if, in the end, his results 



^ A term already applied by him in 1875 to the corresponding 

 nerve in the Batrachia. [Ency. Brit. 9th edition, vol. L, ait. 

 ' Amphibia.") 



